what is the simple meaning of photosynthesis

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Development of the idea

Overall reaction of photosynthesis.

• Basic products of photosynthesis
• Evolution of the process
• Light intensity and temperature
• Carbon dioxide
• Internal factors
• Energy efficiency of photosynthesis
• Structural features
• Light absorption and energy transfer
• The pathway of electrons
• Evidence of two light reactions
• Photosystems I and II
• Quantum requirements
• The process of photosynthesis: the conversion of light energy to ATP
• Elucidation of the carbon pathway
• Carboxylation
• Isomerization/condensation/dismutation
• Phosphorylation
• Regulation of the cycle
• Products of carbon reduction
• Photorespiration
• Carbon fixation in C 4 plants
• Carbon fixation via crassulacean acid metabolism (CAM)
• Differences in carbon fixation pathways
• The molecular biology of photosynthesis

Why is photosynthesis important?

What is the basic formula for photosynthesis, which organisms can photosynthesize.

photosynthesis

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• Roger Williams University Pressbooks - Introduction to Molecular and Cell Biology - Photosynthesis
• BCcampus Open Publishing - Concepts of Biology – 1st Canadian Edition - Overview of Photosynthesis
• photosynthesis - Children's Encyclopedia (Ages 8-11)
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• Table Of Contents

Photosynthesis is critical for the existence of the vast majority of life on Earth. It is the way in which virtually all energy in the biosphere becomes available to living things. As primary producers, photosynthetic organisms form the base of Earth’s food webs and are consumed directly or indirectly by all higher life-forms. Additionally, almost all the oxygen in the atmosphere is due to the process of photosynthesis. If photosynthesis ceased, there would soon be little food or other organic matter on Earth, most organisms would disappear, and Earth’s atmosphere would eventually become nearly devoid of gaseous oxygen.

The process of photosynthesis is commonly written as: 6CO 2 + 6H 2 O → C 6 H 12 O 6 + 6O 2 . This means that the reactants, six carbon dioxide molecules and six water molecules, are converted by light energy captured by chlorophyll (implied by the arrow) into a sugar molecule and six oxygen molecules, the products. The sugar is used by the organism, and the oxygen is released as a by-product.

The ability to photosynthesize is found in both eukaryotic and prokaryotic organisms. The most well-known examples are plants, as all but a very few parasitic or mycoheterotrophic species contain chlorophyll and produce their own food. Algae are the other dominant group of eukaryotic photosynthetic organisms. All algae, which include massive kelps and microscopic diatoms , are important primary producers.  Cyanobacteria and certain sulfur bacteria are photosynthetic prokaryotes, in whom photosynthesis evolved. No animals are thought to be independently capable of photosynthesis, though the emerald green sea slug can temporarily incorporate algae chloroplasts in its body for food production.

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photosynthesis , the process by which green plants and certain other organisms transform light energy into chemical energy . During photosynthesis in green plants, light energy is captured and used to convert water , carbon dioxide , and minerals into oxygen and energy-rich organic compounds .

It would be impossible to overestimate the importance of photosynthesis in the maintenance of life on Earth . If photosynthesis ceased, there would soon be little food or other organic matter on Earth. Most organisms would disappear, and in time Earth’s atmosphere would become nearly devoid of gaseous oxygen. The only organisms able to exist under such conditions would be the chemosynthetic bacteria , which can utilize the chemical energy of certain inorganic compounds and thus are not dependent on the conversion of light energy.

Energy produced by photosynthesis carried out by plants millions of years ago is responsible for the fossil fuels (i.e., coal , oil , and gas ) that power industrial society . In past ages, green plants and small organisms that fed on plants increased faster than they were consumed, and their remains were deposited in Earth’s crust by sedimentation and other geological processes. There, protected from oxidation , these organic remains were slowly converted to fossil fuels. These fuels not only provide much of the energy used in factories, homes, and transportation but also serve as the raw material for plastics and other synthetic products. Unfortunately, modern civilization is using up in a few centuries the excess of photosynthetic production accumulated over millions of years. Consequently, the carbon dioxide that has been removed from the air to make carbohydrates in photosynthesis over millions of years is being returned at an incredibly rapid rate. The carbon dioxide concentration in Earth’s atmosphere is rising the fastest it ever has in Earth’s history, and this phenomenon is expected to have major implications on Earth’s climate .

Requirements for food, materials, and energy in a world where human population is rapidly growing have created a need to increase both the amount of photosynthesis and the efficiency of converting photosynthetic output into products useful to people. One response to those needs—the so-called Green Revolution , begun in the mid-20th century—achieved enormous improvements in agricultural yield through the use of chemical fertilizers , pest and plant- disease control, plant breeding , and mechanized tilling, harvesting, and crop processing. This effort limited severe famines to a few areas of the world despite rapid population growth , but it did not eliminate widespread malnutrition . Moreover, beginning in the early 1990s, the rate at which yields of major crops increased began to decline. This was especially true for rice in Asia. Rising costs associated with sustaining high rates of agricultural production, which required ever-increasing inputs of fertilizers and pesticides and constant development of new plant varieties, also became problematic for farmers in many countries.

A second agricultural revolution , based on plant genetic engineering , was forecast to lead to increases in plant productivity and thereby partially alleviate malnutrition. Since the 1970s, molecular biologists have possessed the means to alter a plant’s genetic material (deoxyribonucleic acid, or DNA ) with the aim of achieving improvements in disease and drought resistance, product yield and quality, frost hardiness, and other desirable properties. However, such traits are inherently complex, and the process of making changes to crop plants through genetic engineering has turned out to be more complicated than anticipated. In the future such genetic engineering may result in improvements in the process of photosynthesis, but by the first decades of the 21st century, it had yet to demonstrate that it could dramatically increase crop yields.

Another intriguing area in the study of photosynthesis has been the discovery that certain animals are able to convert light energy into chemical energy. The emerald green sea slug ( Elysia chlorotica ), for example, acquires genes and chloroplasts from Vaucheria litorea , an alga it consumes, giving it a limited ability to produce chlorophyll . When enough chloroplasts are assimilated , the slug may forgo the ingestion of food. The pea aphid ( Acyrthosiphon pisum ) can harness light to manufacture the energy-rich compound adenosine triphosphate (ATP); this ability has been linked to the aphid’s manufacture of carotenoid pigments.

General characteristics

The study of photosynthesis began in 1771 with observations made by the English clergyman and scientist Joseph Priestley . Priestley had burned a candle in a closed container until the air within the container could no longer support combustion . He then placed a sprig of mint plant in the container and discovered that after several days the mint had produced some substance (later recognized as oxygen) that enabled the confined air to again support combustion. In 1779 the Dutch physician Jan Ingenhousz expanded upon Priestley’s work, showing that the plant had to be exposed to light if the combustible substance (i.e., oxygen) was to be restored. He also demonstrated that this process required the presence of the green tissues of the plant.

In 1782 it was demonstrated that the combustion-supporting gas (oxygen) was formed at the expense of another gas, or “fixed air,” which had been identified the year before as carbon dioxide. Gas-exchange experiments in 1804 showed that the gain in weight of a plant grown in a carefully weighed pot resulted from the uptake of carbon, which came entirely from absorbed carbon dioxide, and water taken up by plant roots; the balance is oxygen, released back to the atmosphere. Almost half a century passed before the concept of chemical energy had developed sufficiently to permit the discovery (in 1845) that light energy from the sun is stored as chemical energy in products formed during photosynthesis.

This equation is merely a summary statement, for the process of photosynthesis actually involves numerous reactions catalyzed by enzymes (organic catalysts ). These reactions occur in two stages: the “light” stage, consisting of photochemical (i.e., light-capturing) reactions; and the “dark” stage, comprising chemical reactions controlled by enzymes . During the first stage, the energy of light is absorbed and used to drive a series of electron transfers, resulting in the synthesis of ATP and the electron-donor-reduced nicotine adenine dinucleotide phosphate (NADPH). During the dark stage, the ATP and NADPH formed in the light-capturing reactions are used to reduce carbon dioxide to organic carbon compounds. This assimilation of inorganic carbon into organic compounds is called carbon fixation.

Van Niel’s proposal was important because the popular (but incorrect) theory had been that oxygen was removed from carbon dioxide (rather than hydrogen from water, releasing oxygen) and that carbon then combined with water to form carbohydrate (rather than the hydrogen from water combining with CO 2 to form CH 2 O).

By 1940 chemists were using heavy isotopes to follow the reactions of photosynthesis. Water marked with an isotope of oxygen ( 18 O) was used in early experiments. Plants that photosynthesized in the presence of water containing H 2 18 O produced oxygen gas containing 18 O; those that photosynthesized in the presence of normal water produced normal oxygen gas. These results provided definitive support for van Niel’s theory that the oxygen gas produced during photosynthesis is derived from water.

ENCYCLOPEDIC ENTRY

Photosynthesis.

Photosynthesis is the process by which plants use sunlight, water, and carbon dioxide to create oxygen and energy in the form of sugar.

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• Photosynthesis (Google doc)

Most life on Earth depends on photosynthesis .The process is carried out by plants, algae, and some types of bacteria, which capture energy from sunlight to produce oxygen (O 2 ) and chemical energy stored in glucose (a sugar). Herbivores then obtain this energy by eating plants, and carnivores obtain it by eating herbivores.

The process

During photosynthesis, plants take in carbon dioxide (CO 2 ) and water (H 2 O) from the air and soil. Within the plant cell, the water is oxidized, meaning it loses electrons, while the carbon dioxide is reduced, meaning it gains electrons. This transforms the water into oxygen and the carbon dioxide into glucose. The plant then releases the oxygen back into the air, and stores energy within the glucose molecules.

Chlorophyll

Inside the plant cell are small organelles called chloroplasts , which store the energy of sunlight. Within the thylakoid membranes of the chloroplast is a light-absorbing pigment called chlorophyll , which is responsible for giving the plant its green color. During photosynthesis , chlorophyll absorbs energy from blue- and red-light waves, and reflects green-light waves, making the plant appear green.

Light-dependent Reactions vs. Light-independent Reactions

While there are many steps behind the process of photosynthesis, it can be broken down into two major stages: light-dependent reactions and light-independent reactions. The light-dependent reaction takes place within the thylakoid membrane and requires a steady stream of sunlight, hence the name light- dependent reaction. The chlorophyll absorbs energy from the light waves, which is converted into chemical energy in the form of the molecules ATP and NADPH . The light-independent stage, also known as the Calvin cycle , takes place in the stroma , the space between the thylakoid membranes and the chloroplast membranes, and does not require light, hence the name light- independent reaction. During this stage, energy from the ATP and NADPH molecules is used to assemble carbohydrate molecules, like glucose, from carbon dioxide.

C3 and C4 Photosynthesis

Not all forms of photosynthesis are created equal, however. There are different types of photosynthesis, including C3 photosynthesis and C4 photosynthesis. C3 photosynthesis is used by the majority of plants. It involves producing a three-carbon compound called 3-phosphoglyceric acid during the Calvin Cycle, which goes on to become glucose. C4 photosynthesis, on the other hand, produces a four-carbon intermediate compound, which splits into carbon dioxide and a three-carbon compound during the Calvin Cycle. A benefit of C4 photosynthesis is that by producing higher levels of carbon, it allows plants to thrive in environments without much light or water. The National Geographic Society is making this content available under a Creative Commons CC-BY-NC-SA license . The License excludes the National Geographic Logo (meaning the words National Geographic + the Yellow Border Logo) and any images that are included as part of each content piece. For clarity the Logo and images may not be removed, altered, or changed in any way.

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Photosynthesis

Reviewed by: BD Editors

Photosynthesis Definition

Photosynthesis is the biochemical pathway which converts the energy of light into the bonds of glucose molecules. The process of photosynthesis occurs in two steps. In the first step, energy from light is stored in the bonds of adenosine triphosphate (ATP), and nicotinamide adenine dinucleotide phosphate (NADPH). These two energy-storing cofactors are then used in the second step of photosynthesis to produce organic molecules by combining carbon molecules derived from carbon dioxide (CO 2 ). The second step of photosynthesis is known as the Calvin Cycle. These organic molecules can then be used by mitochondria to produce ATP, or they can be combined to form glucose, sucrose, and other carbohydrates. The chemical equation for the entire process can be seen below.

Photosynthesis Equation

Above is the overall reaction for photosynthesis. Using the energy from light and the hydrogens and electrons from water, the plant combines the carbons found in carbon dioxide into more complex molecules. While a 3-carbon molecule is the direct result of photosynthesis, glucose is simply two of these molecules combined and is often represented as the direct result of photosynthesis due to glucose being a foundational molecule in many cellular systems. You will also notice that 6 gaseous oxygen molecules are produced, as a by-produce. The plant can use this oxygen in its mitochondria during oxidative phosphorylation . While some of the oxygen is used for this purpose, a large portion is expelled into the atmosphere and allows us to breathe and undergo our own oxidative phosphorylation, on sugar molecules derived from plants. You will also notice that this equation shows water on both sides. That is because 12 water molecules are split during the light reactions, while 6 new molecules are produced during and after the Calvin cycle. While this is the general equation for the entire process, there are many individual reactions which contribute to this pathway.

Stages of Photosynthesis

The light reactions.

The light reactions happen in the thylakoid membranes of the chloroplasts of plant cells. The thylakoids have densely packed protein and enzyme clusters known as photosystems . There are two of these systems, which work in conjunction with each other to remove electrons and hydrogens from water and transfer them to the cofactors ADP and NADP + . These photosystems were named in the order of which they were discovered, which is opposite of how electrons flow through them. As seen in the image below, electrons excited by light energy flow first through photosystem II (PSII), and then through photosystem I (PSI) as they create NADPH. ATP is created by the protein ATP synthase , which uses the build-up of hydrogen atoms to drive the addition of phosphate groups to ADP.

The entire system works as follows. A photosystem is comprised of various proteins that surround and connect a series of pigment molecules . Pigments are molecules that absorb various photons, allowing their electrons to become excited. Chlorophyll a is the main pigment used in these systems, and collects the final energy transfer before releasing an electron. Photosystem II starts this process of electrons by using the light energy to split a water molecule, which releases the hydrogen while siphoning off the electrons. The electrons are then passed through plastoquinone, an enzyme complex that releases more hydrogens into the thylakoid space . The electrons then flow through a cytochrome complex and plastocyanin to reach photosystem I. These three complexes form an electron transport chain , much like the one seen in mitochondria. Photosystem I then uses these electrons to drive the reduction of NADP + to NADPH. The additional ATP made during the light reactions comes from ATP synthase, which uses the large gradient of hydrogen molecules to drive the formation of ATP.

The Calvin Cycle

With its electron carriers NADPH and ATP all loaded up with electrons, the plant is now ready to create storable energy. This happens during the Calvin Cycle , which is very similar to the citric acid cycle seen in mitochondria. However, the citric acid cycle creates ATP other electron carriers from 3-carbon molecules, while the Calvin cycle produces these products with the use of NADPH and ATP. The cycle has 3 phases, as seen in the graphic below.

During the first phase, a carbon is added to a 5-carbon sugar, creating an unstable 6-carbon sugar. In phase two, this sugar is reduced into two stable 3-carbon sugar molecules. Some of these molecules can be used in other metabolic pathways, and are exported. The rest remain to continue cycling through the Calvin cycle. During the third phase, the five-carbon sugar is regenerated to start the process over again. The Calvin cycle occurs in the stroma of a chloroplast. While not considered part of the Calvin cycle, these products can be used to create a variety of sugars and structural molecules.

Products of Photosynthesis

The direct products of the light reactions and the Calvin cycle are 3-phosphoglycerate and G3P, two different forms of a 3-carbon sugar molecule. Two of these molecules combined equals one glucose molecule, the product seen in the photosynthesis equation. While this is the main food source for plants and animals, these 3-carbon skeletons can be combined into many different forms. A structural form worth note is cellulose , and extremely strong fibrous material made essentially of strings of glucose. Besides sugars and sugar-based molecules, oxygen is the other main product of photosynthesis. Oxygen created from photosynthesis fuels every respiring organism on the planet.

Lodish, H., Berk, A., Kaiser, C. A., Krieger, M., Scott, M. P., Bretscher, A., . . . Matsudaira, P. (2008). Molecular Cell Biology 6th. ed . New York: W.H. Freeman and Company. Nelson, D. L., & Cox, M. M. (2008). Principles of Biochemistry . New York: W.H. Freeman and Company.

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Photosynthesis

Photosynthesis n., plural: photosyntheses [ˌfŏʊ.ɾoʊ.ˈsɪn̪.θə.sɪs] Definition: the conversion of light energy into chemical energy by photolithorophs

Table of Contents

Photosynthesis is a physio-chemical process carried out by photo-auto-lithotrophs by converting light energy into chemical energy . Among the endless diversity of living organisms in the world, producers are a unique breed.

Unlike consumers ( herbivores , carnivores , omnivores , or decomposers ) that rely upon other living organisms for their nutritional requirements and nourishment, producers have been distinguished by their ability to synthesize their own food. This is the reason that we call producers “autotrophic or self-reliable” in nature while consumers of all the different categories are called “heterotrophic or dependent” in nature.

Now among producers, there are different categories of producers, i.e. different mechanisms via which they produce their own food.

• Photo-auto-litho-trophs: Since these organisms tend to derive their nutrition by channeling the sun’s light energy, they are termed phototrophic in nature. Also, since they utilize inorganic carbon and translate it into organic carbon atoms, i.e. their means of deriving food becomes autotrophic. Additionally, since the source of electrons (electron donors) here are inorganic compounds, they are specified as lithotrophic . In totality, they can be called photo-auto-litho-trophic in nature. Example : Green plants are nature’s brilliant entities that come under this category. They carry out a photosynthesis cycle by taking in carbon dioxide and fixing it into carbohydrates (energy storage molecule). Some of them also give out oxygen gas that’s vital for the other life forms to survive in the earth’s atmosphere.
• Chemo-auto-lithotrophs: Many of us might be unaware of the fact that there are some autotrophs that don’t utilize sunlight. Rather they derive their energy stored from a different energy source like oxidation of inorganic compounds.

The scope of today’s discussion is limited to photosynthesis and photoautotrophs. So, let’s get started and get to know the answers to these common questions: what is the photosynthesis process, what are the 3 stages of photosynthesis, what does photosynthesis produce, what is a byproduct of photosynthesis, what is the purpose of photosynthesis, is photosynthesis a chemical change, the various inputs and outputs of photosynthesis, which organisms perform photosynthesis , and many other more questions!!!

What is Photosynthesis?

Photosynthesis definition: Photosynthesis is a physio-chemical process carried out by photo-auto-lithotrophs . In simpler language, photosynthesis is the process by which green plants convert light energy into ‘chemical energy’.

This energy transformation is only possible due to the presence of the miraculous pigment molecule chlorophyll in photosynthesis. The chemical energy as referred to before is the fixed carbon molecules generated during photosynthesis.

Green plants and algae have the ability to utilize carbon dioxide molecules and water and produce food (carbohydrates) for all life forms on Earth. There’s no doubt in the fact that life is impossible and unimaginable without green plants that photosynthesize and sustain the cycles of life.

Let’s give you a brief outline of the topic before we head forward.

• Etymology: The photosynthesis process finds its origin in 2 Greek words, firsts one being “phōs (φῶς)” meaning ‘light’ and the second one being “sunthesis (σύνθεσις)” meaning ‘putting together’ . The process of photosynthesis aids the conversion of light energy to chemical energy in varied forms of carbohydrate molecules like sugar molecules and starches.
• Organisms that perform photosynthesis: The organisms are called photo-auto-litho-trophs or simply photoautotrophs.
• Atmospheric gas consumed: Photosynthesizing organisms utilize carbon dioxide in photosynthesis (CO 2 ).
• Atmospheric gas released by “some” photosynthetic organisms (MIND IT-Not all): Some photosynthesizing organisms convert carbon dioxide and aid the process of producing oxygen gas (O2).
• Examples of photosynthesizing organisms: Green plants, cyanobacteria (earlier termed as blue-green algae), and different types of algae that essentially carry out phytoplankton photosynthesis.
• Why is photosynthesis important? The important function of photosynthesis: Food supply for the organisms on Earth, Oxygen supply for the survival of all organisms.
• Site of photosynthesis: Leaves and green tissues. (So when asked where photosynthesis takes place, we can tell that it is this site.)
• What are the reactants of photosynthesis: Carbon dioxide molecules + Water molecules + Light energy
• Products of photosynthesis: Fixed carbon (carbohydrates) + Oxygen (some cases) + Water

Watch this vid about photosynthesis:

Biology Definition: Photosynthesis is the synthesis of complex organic material using carbon dioxide , water , inorganic salts , and light energy (from sunlight) captured by light-absorbing pigments , such as chlorophyll and other accessory pigments . Photosynthesis may basically be simplified via this equation: 6CO 2 +12H 2 O+energy=C 6 H 12 O 6 +6O 2 +6H 2 O, wherein carbon dioxide (CO 2 ), water (H 2 O), and light energy are utilized to synthesize an energy-rich carbohydrate like glucose (C 6 H 12 O 6 ). Other products are water and oxygen .

• Photosynthesis occurs in plastids (e.g. chloroplasts ), which are membrane-bounded organelles containing photosynthetic pigments (e.g. chlorophyll ), within the cells of plants and algae .
• In photosynthetic bacteria ( cyanobacteria ) that do not have membrane-bounded organelles, photosynthesis occurs in the thylakoid membranes in the cytoplasm .

Etymology: from the Greek photo-, “light”, and synthesis, “putting together” Related forms: photosynthetic (adjective) Compare: chemosynthesis See also: photoautotroph

Types of Photosynthesis

Plant photosynthesis and photosynthetic organisms can be classified under different categories on the basis of some characteristic features. They are:

• Types of organisms that carry out photosynthesis on the basis of “cellular structure” Both prokaryotic and eukaryotic organisms carry out photosynthesis.
• Photosynthetic prokaryotes: for example, cyanobacteria
• Eukaryotic: for example, protists ( diatoms , dinoflagellates , Euglena) and green plants. In particular, algae photosynthesis can be observed in green algae , red algae , brown algae , & land plants, like bryophytes , pteridophytes, gymnosperms , and angiosperms .
• Prokaryotic ONLY (anoxygenic photosynthetic bacteria, green sulfur bacteria and purple bacteria)

Photosynthesis: a two-stage process

Photosynthesis is an example of a metabolic process with 2 stages. Both the stages need light (direct or indirect sunlight). Hence, the long-claimed notion of the 2 processes being ‘absolute LIGHT and DARK reactions’ isn’t apt.

Scientific studies have pointed out that even the 2nd stage of photosynthesis requires indirect sunlight. Therefore, rather than classifying the stages as light and dark photosynthesis reactions, we’ll like to classify the 2 stages as follows:

• Photochemical Reaction Process: Light energy is converted to ATP ; photophosphorylation process (light-dependent reactions)
• Through Calvin cycle: In oxygenic photosynthesis as well as anoxygenic photosynthesis
• Through Non-Calvin cycle: Only is some anoxygenic photosynthesis

Evolution of Photosynthesis Process

It is postulated that the very first photosynthetic beings and photosynthesis evolved quite early down the evolutionary timescale of life.

It is also believed that the first photosynthetic beings would have initially resorted to other available reducing agents like hydrogen ions or hydrogen sulfide in contrast to the modern-day photosynthetic organisms that utilize water as the “prime and only sources of electrons”.

It is believed that cyanobacteria would have appeared on the surface of Earth much later than the first photosynthetic beings. Once appeared they must have saturated the Earth’s atmosphere with oxygen gas and led to its oxygenation. Only after the Earth was oxygenated, the more complex forms of life would have later evolved.

When we compare photosynthesis to other metabolic processes like respiration, we can clearly notice that these two processes are almost opposite to each other. But another point to note is that both the processes in synchrony sustain life on Earth.

You cannot separate respiration from photosynthesis or photosynthesis from respiration and expect life to run normally. It is not possible that way. Let’s try to compare and list some characteristic features of photosynthesis and cellular respiration processes.

Photosynthesis vs. Respiration

• Photosynthesis: Anabolic process
• Cellular respiration: Catabolic process By anabolic, we mean the photosynthesis process “utilizes energy to build biomolecules” like carbohydrates, starch, and sugars. These biomolecules are further utilized by both the plants and the organisms dependent on plants for their nutritional needs. On the other hand, respiration is a catabolic process. This energy is utilized to break down complex molecules to derive nutrition out of them.
• Photosynthesis: In the chloroplasts of the eukaryotic phototrophic cells.
• Respiration: Primarily in the mitochondria of the cell.
• Photosynthesis: Carbon dioxide molecules + Water molecules + Light energy
• Respiration: Glucose + Oxygen
• Photosynthesis: Fixed carbon (carbohydrates) + Oxygen (some cases) + Water
• Respiration: Carbon dioxide + Water +energy (ATP)
• Photosynthesis: Endergonic and endothermic
• Respiration: Exergonic and exothermic Just note that these terms endergonic and endothermic both convey the same meaning of “absorbing heat”. And the terms exergonic and exothermic also convey the same meaning of “releasing heat”. The only difference is that –gonics relates to “the relative change in the free energy of the system” while –thermic relates to “the relative change in enthalpy of the system”.
• Photosynthesis: 6CO 2 + 6H 2 O → C6H 12 O 6 + 6O 2
• Respiration: C 6 H 12 O 6 6 + 6O 2 → 6CO 2 + 6H 2 O

Photosynthetic Membranes and Organelles

When we begin the discussion on this topic, it’s important that we know that no photosynthesis is possible without the pigment molecules that absorb light. The absorption of sunlight is the most vital step of photosynthesis.

We should also note that the energy of photons is different for every light of different wavelengths. And the energy needed for the photosynthesis to be conducted is of “a very specific wavelength range”.

For the absorption of lights of desired wavelengths, phototrophs organize their pigment molecules in the form of reaction center proteins . These proteins are located in the membranes of the organisms. Let’s learn how these pigment molecules reside inside the organism and how they make the membranes photosynthetic in nature.

• Prokaryotic photosynthetic organisms: These organisms have their pigment systems or photosystems located in the cell membranes or the thylakoid membranes in the cytosol itself. There are no special organelles called chloroplasts in the prokaryotes.

• Eukaryotic photosynthetic organisms (like green plants): These organisms have their pigment systems or photosystems located in the thylakoids of the chloroplast membranes. Eukaryotes have specialized organelles called chloroplasts (chlorophyll-containing plastids) in their cells.

Photosynthetic Pigments

There are 2 types of photosynthetic pigments in the oxygenic photosynthesizing organisms . They are as follows:

• Porphyrin-derivatives (Chlorophyll in plants and Phycobilin)

Carotenoids

Chlorophyll.

Chlorophyll is the green-colored pigment essential for photosynthesis. Let’s try to list its major characteristic features and roles of it.

• Nature: Lipid
• Location: Embedded in the thylakoid membrane
• Types: 9 types as identified by Arnoff and Allen in 1966 (chlorophyll-a, b, c, d, e, bacteriochlorophyll a, b, chlorobium chlorophyll-650,666). Bacteriochlorophylls are present in the anoxygenic photosynthetic organisms.
• Primary photosynthetic pigment: Chlorophyll-a
• Presence: In all oxygenic photosynthetic organisms
• Absorption range: Visible (blue and red) and IR (Infra-red)
• Ion important for its biological functioning: Magnesium ion (Mg 2+ )
• Structure: Chlorophyll-a, b, and d are “ chlorin ” derivatives; c is a “ porphyrin ” derivative.
• Chlorophyll Tail: Oxygenic photosynthetic organisms have a “ phytol ” tail in their chlorophyll; anoxygenic photosynthetic organisms have a “ geranyl ” tail in their bacteriochlorophylls.
• Main pigment for capturing and storing solar energy
• Photochemical reaction (chlorophyll-a is present in the photochemical reaction center i.e. PCRC. Chlorophyll a, b, c, and d play a role in resonance energy transfer.)

Carotenoid is the photosynthetic pigment essential for working in conjunction with chlorophyll. Let’s try to list its major characteristic features and roles of it.

• Nature: Lipid-soluble
• Types: More than 150
• Absorption range: 400-500nm
• Forms: Carotene (simple hydrocarbon, for example, beta carotene) and xanthophyll (oxygenated hydrocarbon, for example, lutein)
• In excitation and resonance energy transfer
• Photo-protection (work as a free-radical scavenger as well as a quencher)

Phycobilins

Phycobilins aren’t present in all the oxygenic photosynthetic organisms. They have a tetrapyrrole structure (no need for magnesium ion).

• Types: Phycoerythrobilin, Phycocyanobilins, Allophycocyanobilins When these pigment molecules combine with a water-soluble protein, they form the pigment-protein complex (phycobiliproteins, like phycoerythrin and phycocyanin).
• Location: Since these phycobiliproteins are water-soluble, they can’t exist in the membranes like chlorophyll and carotenoids. Therefore, phycobilin pigments as their pigment-protein complex aggregate into clusters and adhere to the membrane. These clusters are called phycobilisomes .
• Exceptional Note: These are the only pigments that are associated with protein molecules.
• Role: Resonance energy transfer

Organelle for Photosynthesis

What is chloroplast? In eukaryotes, photosynthesis occurs in chloroplasts as they are the designated organelles for the photosynthesis process. There are nearly 10-100 chloroplasts in a typical plant cell .

Inside chloroplasts are the thylakoids; the very specific site for the light capturing. The structure of this very unique part of the chloroplasts is briefly discussed here.

Thylakoid is a membrane-bound compartment in the chloroplasts of eukaryotic organisms. They are also present as such in the cytosol of cyanobacteria (cyanobacteria don’t have chloroplasts but they have simply thylakoids).

These thylakoids are the “primary site of the 1st stage of photosynthesis. i.e. “photochemical reaction” or popularly called “light-dependent reactions of photosynthesis”. The main components of the thylakoid are membrane, lumen, and lamellae. The chlorophyll molecules are present inside these thylakoid membranes.

Light-dependent Reactions

The first stage of photosynthesis is popularly called “light-dependent reactions” . We choose to call this stage the “1st stage: PHOTOCHEMICAL REACTION STAGE”. It is also called the “thylakoid reaction stage” or “hill’s reaction” .

This stage is marked by 3 essential steps of photosynthesis: Oxidation of water , reduction of NADP + , and ATP formation . The site where these reactions occur is the lamellar part of the chloroplast. The units of light-dependent reactions are quantosomes .

Let’s discuss this stage under some subheadings:

Wavelengths of light involved and their absorption

The white light that reaches Earth has subparts of different wavelengths together constituting the visible spectrum (390-760nm). But the photosynthetic organisms specifically use a subpart called PAR ( P hotosynthetically A ctive R adiation).

PAR ranges from 400-760nm. Blue light is 470-500nm while red light is 660-760nm). The green light (500-580nm) is reflected back by the plants and this is the reason that plants appear green in color. Blue-green light is not used, only blue light is used.

Absorption spectrum and action spectrum

• Absorption Spectrum: This is a pigment-specific entity or terminology. To find the absorption spectrum of a pigment, you need to plot “the amount of absorption of different wavelengths of light by that particular pigment” . The graph has the “wavelengths of light (in nanometers/nm)” on the X-axis and the “percentage of light absorption” on the Y-axis.

• Action Spectrum: To find the action spectrum of a pigment, you need to plot the “effectiveness of the different wavelengths of light in stimulating photosynthesis process” . The graph has the “wavelengths of light (in nanometers/nm)” on the X-axis and the “rate of photosynthesis (measured as oxygen released)” on the Y-axis. When you superimpose the action spectrum of photosynthesis with the absorption spectrum of the specific pigment, you can find the contribution of each different wavelength in the photosynthesis rate, photosynthetic efficiency, and photosynthetic productivity.

IMPORTANT NOTE: The absorption spectrum is calculated for any of the many pigments involved in photosynthesis. Contrastingly, the action spectrum is calculated only for the photochemical reaction performing pigment i.e. chlorophyll-a present at the reaction center. We identify the progress of photochemical reactions as the “evolution of oxygen gas” that primarily happens at the reaction center where only chlorophyll-a is present. Since the action is directly correlated to the specific excitation of chlorophyll-a molecule, the action spectrum is scientifically calculated only for this chlorophyll-a.

• Absorption spectrum of chlorophyll- a : 430 nm (blue), 660nm (red) {more absorbance at 660 nm)
• Absorption spectrum of chlorophyll-b: 430 nm (blue), 660nm (red) {more absorbance at 430 nm)

What actually happens in the Light-dependent reaction

Let’s briefly describe what actually happens here.

• 1 photon is absorbed by 1 molecule of the chlorophyll (P680) and simultaneously 1 electron is lost here.
• The electron flow of the photochemical reaction begins here.
• The electron is transferred to D1/D2 protein, then to a modified form of chlorophyll and “pheophytin”.
• After that, it’s transferred to plastoquinone A and then B.
• Initiates an electron flow down an electron transport chain.
• Ultimately aids the NADP reduction to NADPH.
• Creation of a proton gradient across the chloroplast membrane.
• Further on this proton gradient is exploited by the ATP synthase for the generation of ATP molecules.

Water photolysis

Now, if you are wondering how the first electron lost by the 1st chlorophyll is replenished to keep this cycle going, read on. The answer to this query is “photolysis of water molecules” . The chlorophyll molecule regains the lost electron when the “oxygen-evolving complex” in the thylakoid membrane carries out the photolysis of water. The chlorophyll molecule ultimately regains the electron it lost when a water molecule is split in a process called photolysis, which releases oxygen.

Many scientists had a doubt about the source of oxygen in photosynthesis. Some speculated the oxygen atom of the CO 2 gas is the source of oxygen post-photosynthesis. But it was the collective contribution of some 4 scientists that gave clarity on this topic.

C.B. Van Niel worked on purple photosynthetic bacteria ( Chromatium vinosum ) and found out that the source of oxygen is the oxidation of water molecules (‘indirect evidence’). While Ruben, Hassid, and Kamen carried out an isotopic study that gave ‘direct evidence’ of oxygen-evolving from H 2 O molecules and not CO 2 molecules.

Hydrolysis of 2 molecules of water leads to the evolution of 1 molecule of oxygen gas. The photosynthesis equation for light-dependent reactions (non-cyclic electron flow) or the chemical formula for photosynthesis:

2 H 2 O + 2 NADP+ + 3 ADP + 3 Pi + light → 2 NADPH + 2 H+ + 3 ATP + O 2

The photochemical reaction (or the light-dependent reactions) can be classified as:

• Cyclic reaction: Only 1 photosystem ( PS1 ) is involved. (Photon excites P700 in PS1, electron reaches Fe-S, then Ferredoxin, then Plastoquinone and then Cyt b6f complex and then Plastocyanin). Since in the solo involvement of PS1 here, the electron flow becomes cyclic. And this phosphorylation process is called cyclic phosphorylation. It happens in the stroma lamellae when light beyond 680nm is available.
• Non-cyclic reaction: Both photosystems (PS1 and PS2 ) are involved. (Photon excites P680 in PS2, the electron is lost and transferred to pheophytin, then sent on a roller coaster (Z-scheme). Within the z-scheme, the final redox reaction enables the reduction of NADP+ to NADPH. And the chemiosmotic potential generation via proton pumping proton across the membrane and into the thylakoid lumen ensures ATP synthesis.

Data Source: Akanksha Saxena of Biology Online

Light-Independent Reactions (Carbon-fixation Reaction)

Also called the carbon fixation process, the “light-independent reactions” is a misnomer as Science has now already proved that the second stage of photosynthesis isn’t really light-independent reactions. Though it doesn’t need direct light, indirect light is involved even in this process. We choose to label this stage of photosynthesis as the “2nd stage: CARBON-FIXATION REACTION STAGE ”, which is also called:

• Calvin Cycle or “stromal reaction” as it manifests in the stroma part of the chloroplast
• “C3 Cycle” or the “reductive pentose phosphate cycle”

Calvin cycle

The inputs for the Calvin cycle  in most plants come from the previously occurred photochemical reaction. In this cycle, the carbon dioxide produced is fixed to a glucose molecule. To be very specific, the Calvin cycle directly doesn’t produce glucose, rather it produces glyceraldehydes-5-phosphate (G-3-P). Glucose is formed after these G-3-P molecules move into the cytosol from the chloroplast .

It consists of primarily 3 steps as follows:

• Carboxylation: Acceptance of CO 2 by RuBP which is a 5-carbon compound and the CO2-acceptor). 2 molecules of 3-phosphoglycerate are generated as the result of the carboxylation process.
• Reduction: Generation of 3C/4C/5C/6C/7C molecules.
• Regeneration of RUBP: 3 molecules of RuBP are regenerated.

In totality, 3 molecules of CO 2 produce 1 molecule of G-3-P. This uses 9 ATPs and 6 NADPHs. And, 6 molecules of CO 2 produce 2 molecules of G-3-P which further produce 1 molecule of glucose. This uses 18ATPs and 12 NADPHs.

The main enzyme is RuBisCo . It’s a multi-enzyme complex with 8 large and 8 small subunits. The substrates for this enzyme are CO 2 , O 2 , and RuBP. An essential ion for the biological functioning of this enzyme: Mg 2+ . The role of RuBisCo is that it captures carbon dioxide gas from the atmosphere and utilizes the NADPH from the 1st stage (photochemical reaction/light-dependent reaction stage) to fix the CO 2 .

The equation of dark reaction of photosynthesis/light-independent reaction stage/2nd stage is: 3 CO 2 + 9 ATP + 6 NADPH + 6 H + → C 3 H 6 O 3 -phosphate + 9 ADP + 8 Pi + 6 NADP+ + 3 H 2 O

The simple carbon sugars formed via the C3 cycle are utilized by the biological systems to form complex organic compounds like cellulose, precursors for amino acids synthesis and thereby proteins, precursors for lipids, and the source of fuel for respiration.

Important Point To Note: It happens in all the photosynthetic organisms as the basic carbon-fixation step.

Carbon concentrating mechanisms

There are many carbon concentrating mechanisms to increase the carbon dioxide levels and the carbon fixation process like C4, CAM, etc.

• Doesn’t happen in all photosynthetic organisms. Rather it happens in conjunction with the C3 cycle in some 4% of angiosperm families.
• Most commonly angiosperm families that witness C4 cycle: Poaceae, Cyperaceae.
• First explained by: Hatch and Slack (hence also called the Hatch and Slack cycle). They worked on the maize plant.
• Role: Endow the ability to efficiently conduct photosynthesis in plants of the semi-arid regions by making them well adapted.
• Mechanism: By separation of photosynthesis stages in 2 types of cells (mesophyll cells and bundle sheath cells). The light reaction is restricted to the mesophyll cells and the CO 2 fixation happens in the bundle sheath cells. This phenomenon is also termed as “chloroplast dimorphism” in C4 plants. The Kranz anatomy is visible here.
• Why does the need arise in the first place? – In semi-arid regions or regions with very hot and dry environmental conditions, plants are forced to close their stomata in order to limit water loss. Under such harsh conditions, the intake of CO 2 decreases during the day as the stomata are forced closed. This might lead to no CO 2 intake and hence no CO 2 fixation (2nd stage of photosynthesis). But the 1st stage of photosynthesis keeps running as it doesn’t depend on stomata opening or closure. This means that a continuous oxygen evolution happens which can lead to oxygen saturation. As we know that RuBisCo enzymes use O 2 gas as substrate too, and this can lead to an increased rate of photorespiration by the oxygenase activity of RuBisCo. This further decreases the carbon fixation. This is a very big issue if not resolved. Hence, for situations like these, carbon concentrating mechanisms have evolved in some families of plants to concentrate and enrich the CO 2 concentration in the leaves of these plants under such conditions.
• Important enzyme for CO 2 concentration: PEP carboxylase
• CO 2 is first added to a three-carbon compound called phosphoenolpyruvate (PEP) in this cycle. This leads to the formation of a four-carbon (4C)  molecule called oxaloacetic acid or malate. This step happens in the mesophyll cells of the leaves.
• After that, these 4C compounds are transferred to the bundle sheath cells where the normal C3 cycle fixes them into glucose molecules.
• This CO 2 concentrating mechanism works on the “principle of separating the RuBisCo enzyme from the O 2 -generating photochemical reactions” in order to reduce the rates of photorespiration and simultaneously increase the rates of CO 2 fixation.
• This increases the photosynthetic capacity of the leaf/leaf photosynthesis.
• When the high light and high-temperature conditions are dominant, C4 plants prove more photosynthetically efficient than C3 plants as they produce more sugar molecules in such conditions.
• Examples of C4 plants: Many crop plants like wheat, maize, rice, sorghum, millet, and sugarcane.
• Number of ATPs required: 12 (for C-enrichment) + 18 (for C-fixation)= 30 ATPS for 1 glucose production
• Number of NADPH required: 18 NADPH for 1 glucose production
• Some plants resort to another mechanism called the CAM cycle in conjunction with the C3 cycle to fix carbon dioxide.
• Examples: xerophytes like cactus photosynthesis, and most succulents.
• Around 16,000 species of plants utilize the CAM mechanism
• Mechanism: Utilize PEP carboxylase to capture carbon dioxide. In contrast to the C4 cycle where there is a “spatial separation of the 2 processes of CO 2 reduction to PEP and PEP fixation to glucose”, CAM plants display a “temporal separation of the 2 listed processes”.

Land plants display different types of photosynthesis based on their requirements and environmental constraints. They are C3, C4 +C3, and CAM+ C3 types of photosynthesis.

Aquatic plants and algae display some extra features in the photosynthetic machinery. These features further refine and define the smooth functioning and efficiency of photosynthesis.

Example: Cyanobacteria photosynthesis – cyanobacteria have carboxysomes  that help in enriching the concentration of carbon dioxide around the RuBisCO enzyme. This directly increases the photosynthetic rates. The distinguished and specially enabled enzyme in the carboxysomes is called “carbonic anhydrase”. The carbonic anhydrase possesses the ability to evolve and release CO 2 from the dissolved hydrocarbonate ions (HCO-). As soon as the CO 2 is released, RuBisCo takes care that it doesn’t go to waste.

Order and Kinetics

There are innumerable reactions and processes involved in the biological mechanism of photosynthesis. Besides the normal flow of photosynthesis, there are some plant-specific and condition-specific additional steps that further complicate the mechanism.

Since every biological mechanism has a lot of enzymes, factors, cofactors, substrates, and entities involved, photosynthesis is no different.

Let’s try to list some kinetics-specific pointers that may help.

As discussed in the overview and starting of this article, the early photosynthetic organisms must have been primarily “anoxygenic” in nature. These bacteria used some other source than water molecules as their primary electron donors. Even the geological evidence aligns with this fact as the early atmosphere of Earth was highly reducing in nature. Some speculated organisms of the early evolutionary phase are :

• Green sulfur bacteria (Electron donor= hydrogen and sulfur)
• Purple sulfur bacteria (Electron donor= hydrogen and sulfur)
• Green nonsulfur bacteria (Electron donor= various amino and other organic acids)
• Purple nonsulfur bacteria (Electron donor= variety of nonspecific organic molecules)

After this, some filamentous photosynthetic organisms are expected to have evolved. This is scaled to be an occurrence of some 3.4 billion years old timeline. It is around 2 million years ago that oxygenic photosynthesis is believed to have evolved.

The modern and more commonly known photosynthesis in plants and most of the photosynthetic prokaryotes= Oxygenic (Electron donor= Water molecules)

Symbiosis and the origin of chloroplasts

There are some animal groups that have the ability to form and establish symbiotic relationships with photosynthetic organisms. By establishing such a relationship, these organisms can directly rely upon their photosynthetic partner for energy and food requirements. Some examples of such animal groups are:

• Sea anemones
• Marine mollusks (example: Elysia viridis & Elysia chlorotica )
• Fungi photosynthesis (Lichens)

When such symbiotic relationships are established, it’s sometimes observed that some genes of the plant cell’s nucleus get transferred to the animal cell . (Observed in some slugs).

Origin of Chloroplasts

Such symbiosis is popularly claimed to be the source of chloroplast evolution. As we notice many similarities between the photosynthetic bacteria and chloroplasts, the evolution of chloroplasts is often hinted to have occurred from these bacteria. Some of the common features between the 2 are:

• Circular chromosome
• Prokaryotic-type ribosome
• A similar set of proteins in the photosynthetic reaction center

It is for all these commonalities the “ endosymbiotic theory ” had been proposed for the evolution of chloroplasts and mitochondria in the eukaryotic cells. According to the endosymbiotic theory, the early eukaryotic cells are believed to have acquired the photosynthetic bacteria by the process of endocytosis). Those early eukaryotic cells after acquiring the photosynthetic bacteria transformed to be self-sustainable and became the “first plant cells”. (Mitochondria photosynthesis is true, they are associated with respiration!)

Photosynthetic eukaryotic lineages

Photosynthetic eukaryotic lineages include:

• Glaucophytes
• Chlorophytes
• Rhodophytes
• Cryptophytes (some clades)
• Haptophytes (some clades)
• Dinoflagellates & chromerids
• Euglenids—clade Excavata (unicellular)

Cyanobacteria and the evolution of photosynthesis

Almost all the prokaryotes carry out anoxygenic photosynthesis in contrast to cyanobacteria, which perform oxygenic photosynthesis. This ability to carry out oxygenic photosynthesis is speculated to have evolved at least 2450–2320 million years ago. The first photosynthetic cyanobacteria might not have been oxygenic as Earth’s atmosphere had no oxygen then.

This topic still requires more scientific study to bring out conclusive results. From the paleontological evidence, it is claimed that the 1st cyanobacteria evolved around 2000 Ma.

For the initial years of the Earth’s oxygen-rich environment (after the oxygen-evolving mechanism evolved), cyanobacteria are claimed to be the “principal primary producers of oxygen”. Even to date, cyanobacteria have been proven vital for marine ecosystems. They’re the primary producers of oxygen in oceans.

Cyanobacteria also fix nitrogen electrons fixation and play a role in biological nitrogen cycles.

Experimental History

We will list the long experimental history in deciphering the extensive photosynthesis process through the ages.

Discovery, Refinements, and Development of the concept

Find out the discovery, refinements, and development of photosynthesis as summarized in the table below:

C3 : C4 photosynthesis research

Several studies were conducted using isotopes of radioactive elements to identify the various aspects of the photosynthetic process. A number of organisms like Chlorella , Stellaria media, Cladophora, Spirogyra, Rhodopseudomonas , sulfur bacteria, green plants like maize, etc have been used to understand the photosynthesis process over the years. Gas exchange studies, isotopic studies, light spectrum studies, radioactive studies, plant anatomical and physiological studies, studies involving roles of carbon dioxide and water, etc have all together opened the gates for our deeper understanding of this topic.

The 3 main factors that directly affect the photosynthesis process are:

• Light irradiance and wavelength
• Carbon dioxide concentration

Temperature

Although there are many more corollary factors, these 3 are the most important ones.

Light intensity (irradiance), wavelength, and temperature

Light is an essential factor for photosynthesis. It directly affects the rate of it. There are 3 different parameters that we should look into:

• Sciophytes : Grow under “diffuse” light. Example: Oxalis
• Heliophytes: Grow under “direct: light. Example: Dalbergia
• Light quality: PAR as previously discussed is the quality or the fraction of light energy that is ‘photosynthetically active’ in nature. It ranges from 400-700nm in wavelength.
• Duration of light: This parameter doesn’t affect the rate of photosynthesis but affects the total photosynthetic output.

Carbon dioxide levels and photorespiration

Carbon dioxide concentration is the major factor in determining the rate of photosynthesis. There is no carbon-dioxide enriching system in C3 plants like the C4 plants. So, if you increase the concentration of CO 2 in the system, the photosynthetic rate of C3 plants will increase as the CO 2 concentration increases. On the other hand, the photosynthetic yield of the C4 plant won’t increase in such a scenario.

• CO 2 Compensation Point: A stage in CO 2 concentration when there’s no absorption of CO 2 by the illuminated plant part.

Featuring… “The curious case of RuBisCO and PEP Carboxylase”

Imagine an equal concentration (50-50%) of the two isotopes of carbon, C-12 and C-13, in the form of 12CO 2 and 13CO 2 , made available to both C3 and C4 plants. Now, can you tell which isotope of the carbon will be fixed more or less by the two types of photosynthetic organisms? Can you guess if there would be a “preferable” isotope between the two? Do you think C3 plants will fix the 12CO 2 and 13CO 2 equally or unequally? Or do you think the 12CO 2 and 13CO 2 incorporation would have a biased ratio in any of the two (C3/C4 plants)????

The answer to this lies in the major carbon fixing enzyme involved.

• C3 plants: Major C-fixing enzyme is RuBisCo and RuBisCo has a “discriminatory ability” to preferably fix 12CO 2 and not 13CO2. Hence, you will find more 12CO 2 fixed than 13CO 2 in the C3 plants.
• C4 plants: Major C-fixing enzyme is not RuBisCo but PEP Carboxylase . PEP Carboxylase has “no discriminatory ability”. So, you’ll find an almost equal proportion of 12CO 2 and 13CO 2 getting fixed in C4 plants. So, in comparison to C3 plants, the chances of getting 13CO 2 fixed are more in C4 plants.

Choose the best answer. 

Send Your Results (Optional)

• Rutherford, A.W., Faller, P. (Jan 2003). “Photosystem II: evolutionary perspectives”. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 358 (1429): 245–253. doi:10.1098/rstb.2002.1186. PMC 1693113. PMID 12594932.
• Arnon, D.I., Whatley, F.R., Allen, M.B. (1954). “Photosynthesis by isolated chloroplasts. II. Photophosphorylation, the conversion of light into phosphate bond energy”. Journal of the American Chemical Society. 76 (24): 6324–6329. doi:10.1021/ja01653a025.
• Ehrenberg, R. (2017-12-15). “The photosynthesis fix”. Knowable Magazine. Annual Reviews. doi:10.1146/knowable-121917-115502. Retrieved 2018-04-03.
• El-Sharkawy, M.A., Hesketh, J.D. (1965). “Photosynthesis among species in relation to characteristics of leaf anatomy and CO 2 diffusion resistances”. Crop Sci. 5 (6): 517–521. doi:10.2135/cropsci1965.0011183x000500060010x.
• Earl, H., Said Ennahli, S. (2004). “Estimating photosynthetic electron transport via chlorophyll fluorometry without Photosystem II light saturation”. Photosynthesis Research. 82 (2): 177–186. doi:10.1007/s11120-004-1454-3. PMID 16151873. S2CID 291238.

©BiologyOnline.com. Content provided and moderated by Biology Online Editors.

Last updated on July 15th, 2022

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Photosynthesis

What is photosynthesis.

It is the process by which green plants, algae, and certain bacteria convert light energy from the sun into chemical energy that is used to make glucose. The word ‘photosynthesis’ is derived from the Greek word phōs, meaning ‘light’ and synthesis meaning ‘combining together.’

Jan Ingenhousz, the Dutch-born British physician and scientist, discovered the process of photosynthesis.

Where does Photosynthesis Occur

Photosynthesis takes place mainly in the leaves of green plants and also in the stems of herbaceous plants as they also contain chlorophyll. Sometimes it also occurs in roots that contain chlorophyll like in water chestnut and Heart-leaved moonseed. Apart from plants, photosynthesis is also found to occur in blue-green algae.

What Happens During Photosynthesis

It involves a chemical reaction where water, carbon dioxide, chlorophyll, and solar energy are utilized as raw materials (inputs) to produce glucose, oxygen, and water (outputs).

Stages of the Process

Photosynthesis occurs in two stages:

1) The Light-dependent Reaction

• Takes place in the thylakoid membranes of chloroplasts only during the day in the presence of sunlight
• High-energy phosphate molecules adenosine triphosphate ( ATP ) and the reducing agent NADPH are produced with the help of electron transport chain

2) The Light-independent or Dark Reaction ( Calvin cycle )

• Takes place in the stroma of chloroplast in the absence of light that helps to fix carbon
• ATP and NADPH produced in the light reaction are utilized along with carbon dioxide to produce sugar in the form of glucose

Factors Affecting the Rate of Photosynthesis

• Intensity of Light: The higher intensity of light increases the rate of photosynthesis
• Temperature:  Warmer the temperature, higher the rate of photosynthesis. The rate is highest between the temperatures of 25° to 35° C, after which it starts to decrease
• Concentration of Carbon dioxide: Higher concentration of carbon dioxide increases the rate of photosynthesis until it reaches a certain point, beyond which no further effects are found   

Although all the above factors together interact to affect the rate of photosynthesis, each of them individually is also capable of directly influencing the process without the other factors and thus called limiting factors.

Importance of Photosynthesis

It serves two main purposes that are essential to support life on earth:

• Producing food for organisms that depend on others for their nutrition such as humans along with all other animals
• Synthesizing oxygen by replacing carbon dioxide in the atmosphere

Ans. Photosynthesis is an endothermic reaction because it absorbs the heat of the sun to carry out the process.

Ans. The oxygen in photosynthesis comes from splitting the water molecules.

Ans. Chlorophyll is the main light-absorbing pigment in photosynthesis.

Ans. The role of water is to provide oxygen in the form of oxygen gas to the atmosphere.

Ans. Sunlight is the source of energy that drives photosynthesis.

Ans. The easiest way to measure the rate of photosynthesis is to quantify the carbon dioxide or oxygen levels using a data logger. The rate of photosynthesis can also be measured by determining the increase in the plant ’s biomass (weight).

Ans. Photosynthesis is an energy-requiring process occurring only in green plants, algae, and certain bacteria that utilizes carbon dioxide and water to produce food in the form of carbohydrates. In contrast, cellular respiration is an energy-releasing process found in all living organisms where oxygen and glucose are utilized to produce carbon dioxide and water.

Ans. Glucose produced in photosynthesis is used in cellular respiration to make ATP.

Article was last reviewed on Tuesday, April 21, 2020

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Middle school biology

Course: middle school biology   >   unit 3, photosynthesis in organisms.

• Understand: photosynthesis in organisms
• Apply: photosynthesis in organisms

Key points:

• Many organisms make their own food through photosynthesis . Plants, algae, and some unicellular organisms do photosynthesis.
• Photosynthesis is powered by energy from sunlight. This energy is used to rearrange atoms in carbon dioxide and water to make oxygen and sugars.
• Carbon dioxide and water are inputs of photosynthesis. These inputs come from the environment.
• The oxygen is released into the environment.
• The sugars can provide chemical energy . This energy can be used by the organism to live and grow. The sugars can also be used to build the organism’s structures, or they can be stored for later use.
• Photosynthesis is usually carried out in chloroplasts . Chloroplasts are cell structures that contain molecules called chlorophyll . Chlorophyll helps capture light energy.

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Photosynthetic Cells

Cells get nutrients from their environment, but where do those nutrients come from? Virtually all organic material on Earth has been produced by cells that convert energy from the Sun into energy-containing macromolecules. This process, called photosynthesis, is essential to the global carbon cycle and organisms that conduct photosynthesis represent the lowest level in most food chains (Figure 1).

What Is Photosynthesis? Why Is it Important?

Most living things depend on photosynthetic cells to manufacture the complex organic molecules they require as a source of energy. Photosynthetic cells are quite diverse and include cells found in green plants, phytoplankton, and cyanobacteria. During the process of photosynthesis, cells use carbon dioxide and energy from the Sun to make sugar molecules and oxygen. These sugar molecules are the basis for more complex molecules made by the photosynthetic cell, such as glucose. Then, via respiration processes, cells use oxygen and glucose to synthesize energy-rich carrier molecules, such as ATP, and carbon dioxide is produced as a waste product. Therefore, the synthesis of glucose and its breakdown by cells are opposing processes.

However, photosynthesis doesn't just drive the carbon cycle — it also creates the oxygen necessary for respiring organisms. Interestingly, although green plants contribute much of the oxygen in the air we breathe, phytoplankton and cyanobacteria in the world's oceans are thought to produce between one-third and one-half of atmospheric oxygen on Earth.

What Cells and Organelles Are Involved in Photosynthesis?

Chlorophyll A is the major pigment used in photosynthesis, but there are several types of chlorophyll and numerous other pigments that respond to light, including red, brown, and blue pigments. These other pigments may help channel light energy to chlorophyll A or protect the cell from photo-damage. For example, the photosynthetic protists called dinoflagellates, which are responsible for the "red tides" that often prompt warnings against eating shellfish, contain a variety of light-sensitive pigments, including both chlorophyll and the red pigments responsible for their dramatic coloration.

What Are the Steps of Photosynthesis?

Photosynthesis consists of both light-dependent reactions and light-independent reactions . In plants, the so-called "light" reactions occur within the chloroplast thylakoids, where the aforementioned chlorophyll pigments reside. When light energy reaches the pigment molecules, it energizes the electrons within them, and these electrons are shunted to an electron transport chain in the thylakoid membrane. Every step in the electron transport chain then brings each electron to a lower energy state and harnesses its energy by producing ATP and NADPH. Meanwhile, each chlorophyll molecule replaces its lost electron with an electron from water; this process essentially splits water molecules to produce oxygen (Figure 5).

Once the light reactions have occurred, the light-independent or "dark" reactions take place in the chloroplast stroma. During this process, also known as carbon fixation, energy from the ATP and NADPH molecules generated by the light reactions drives a chemical pathway that uses the carbon in carbon dioxide (from the atmosphere) to build a three-carbon sugar called glyceraldehyde-3-phosphate (G3P). Cells then use G3P to build a wide variety of other sugars (such as glucose) and organic molecules. Many of these interconversions occur outside the chloroplast, following the transport of G3P from the stroma. The products of these reactions are then transported to other parts of the cell, including the mitochondria, where they are broken down to make more energy carrier molecules to satisfy the metabolic demands of the cell. In plants, some sugar molecules are stored as sucrose or starch.

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What Is Photosynthesis?

The photosystem, the process.

Photosynthesis is the process used by plants to convert sunlight into chemical energy that can be used to fuel the plants’ growth. The process is fueled by the sun and powered by the chloroplasts in the plants’ leaves. The process begins with the sun’s light energy breaking down water molecules into oxygen and hydrogen. The oxygen is released into the air and the hydrogen is used to power the conversion of carbon dioxide into glucose, which is the plant’s food.

Plants, like every other organism, need energy to live, grow and repair. Unlike heterotrophs –  animals that consume food to synthesize energy – plants or autotrophs are self-reliant — they can make their own food, and therefore energy, by using the resources available in their surroundings. The resources include sunlight, water and carbon dioxide, and this incredible process is called  photosynthesis.

The process is known as photo synthesis because, while water and carbon dioxide are the major ingredients required to cook the food, it is light that ignites the stove, and sunlight is the most abundant light that illuminates the planet.

A photosynthetic organism using the energy of photons ( photo-) , makes its own food (- synthesis ).

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A plant is essentially a highly efficient solar panel. It is replete with molecular structures that have evolved to soak up and absorb as much sunlight as possible.

These structures are in separate compartments of the cell called chloroplasts. Inside the chloroplasts are small towers of discs called grana, each disc is called thylakoid. In the membrane or lining of the thylakoids are a dense grid of various light absorbing molecules, the most notable of them, chlorophyll. The pigments, like any other pigments, absorb light of certain wavelengths and reflect the rest.

This is how pigments or colors are radiated. Majority of plants are green because they (the abundance of chlorophyll) reflect wavelengths that we associate with green while absorbing wavelengths that fall into the red and blue zones. But there are other pigments that absorb different wavelengths in the visible spectrum. Carotenoids are pigments that absorb in the blue to green wavelengths, reflecting orange, yellow and red wavelengths.

This is why leaves take on shades from orange to red during autumn , as they start losing their chlorophyll come at that time of the year. Some deep-sea algae (rhodophyta) are red in color due to a photosynthetic pigment called phycoerythrin and phycocyanin which absorb in blue to green region and reflect back red wavelengths.

These pigments are part of larger protein machines called photosystems. Plants have two photosystems – photosystem I (PSI) and photosystem II (PSII), each having a key chlorophyll-a molecule embedded in it, along with other accessory pigments. These accessory pigments collect light energy and pass it onto the main chlorophyll-a molecule. The photosystems are the main light harvesting machines in the chloroplasts that convert light energy to chemical energy that the cell can use to do build its food. Some algae and photosynthetic bacteria only have single photosystems.

Solar energy is used by the chloroplast to trigger a chemical reaction between the two reactants – water and carbon dioxide.

Also Read: Why Are Plants Red In Color At The Bottom Of The Ocean?

A plant obtains water from its roots through osmosis . From there, the water ascends through the stem and is transported to every part of the plant that requires it for a myriad of processes other than photosynthesis.

To ensure that plant doesn’t dry out due to evaporation, they evolved special pores, called stomata about 400 million years ago. Stomata allowed them to respire, exchanging gases like oxygen and carbon dioxide. However, the crucial development of pores came at the expense of losing water unnecessarily

The stomata inhale carbon dioxide exhaled by animals, which reacts with water in the presence of sunlight to create sugar (glucose), their food. However, the reaction unfolds in two parts.

The first part is called a light-dependent or simply, a light reaction, in which light breaks down water to produce oxygen molecules. These molecules are the same oxygen molecules that we breathe. They are exhaled through the stomata and dispersed into the air. The light energy absorbed by a pigment can be either simply dissipated as heat or be converted into another form of energy. We witness the latter in plants. The light reaction converts solar energy into chemical energy; the reaction also produces ATP (Adenosine Tri-Phosphate) and NADP + (Nicotinamide Adenine Dinucleotide Phosphate), organic compounds that become sources of energy for subsequent metabolic processes.

When light breaks down water at PSII. This releases two protons (H+) and oxygen, and two energized electrons which the chlorophyll-a in PSII accepts. These electrons are passed from PSII to other middle protein structures, like cytochrome bf, to PSI. There these two electrons get energized at PSI due to light energy captured by the photosystem’s chlorophyll-a. These electrons are them funneled to making NADPH from NADP+.

While passing these energized electrons, protons (H+) were taken from outside the thylakoid to its inside. This creates a gradient of H+, more inside than outside. This electrochemical gradient fuels the production of ATP. This happens because of a protein machine called ATP synthase. H+ from inside can return to the outside only by passing through ATP synthase. When H+ pass through ATP synthase, it converts ADP to ATP. This is how light and water create energy for the cell.

One of these processes is the next part of the reaction itself. The two sources of energy fuel the light-independent or dark reaction. The energy breaks down the carbon dioxide molecules and reorganizes the constituents to form a molecule of glucose. The chloroplast then harvests energy by breaking down that glucose, just how mitochondria in animal cells produce energy by breaking down the food they consume. Photosynthesis as a combination of the two reactions can be summarized with this expression:

But if plants can produce ATP through photosynthesis, then why do they need to respire? First, photosynthesis can only happen in the daytime, when there is sun around to provide light. Plants still need energy when there is no light. Second, glucose is important molecule for the cell. It can be broken down or built up to make many other biomolecules like DNA , RNA, proteins, and even fats.

In this manner, we share a deep and indispensable symbiotic relationship with plants. The byproduct, or put more impolitely, the waste product exhaled by plants gives us life, whereas carbon dioxide, the waste product that we exhale, gives plants their life. American biologist and one of my favorite science communicators, Lynn Margulis, called this innocuous act of breathing, spirituality.

However, she believes that “the connection doesn’t stop at the exchange of gases in the atmosphere… The fact that we are connected through space and time shows that life is a unitary phenomenon, no matter how we express the fact.”

Also Read: How Do Plants Excrete?

• PHOTOSYNTHESIS - www2.estrellamountain.edu
• What is Photosynthesis.

Akash Peshin is an Electronic Engineer from the University of Mumbai, India and a science writer at ScienceABC. Enamored with science ever since discovering a picture book about Saturn at the age of 7, he believes that what fundamentally fuels this passion is his curiosity and appetite for wonder.

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What is photosynthesis?

Photosynthesis is the process plants, algae and some bacteria use to turn sunlight, carbon dioxide and water into sugar and oxygen.

• Photosynthetic processes
• Photosynthesis equation
• The carbon exchange
• How do plants absorb sunlight?

How does photosynthesis start?

• Location of photosynthesis

Light-dependent reactions

• The Calvin cycle

Types of photosynthesis

Additional resources.

Photosynthesis is the process used by plants, algae and some bacteria to turn sunlight into energy. The process chemically converts carbon dioxide (CO2) and water into food (sugars) and oxygen . The chemical reaction often relies on a pigment called chlorophyll, which gives plants their green color.  Photosynthesis is also the reason our planet is blanketed in an oxygen-rich atmosphere.

Types of photosynthetic processes

There are two types of photosynthesis: oxygenic and anoxygenic. They both follow very similar principles, but the former is the most common and is seen in plants, algae and cyanobacteria. 

During oxygenic photosynthesis, light energy transfers electrons from water (H2O) taken up by plant roots to CO2 to produce carbohydrates . In this transfer, the CO2 is "reduced," or receives electrons, and the water is "oxidized," or loses electrons. Oxygen is produced along with carbohydrates.

This process creates a balance on Earth, in which the carbon dioxide produced by breathing organisms as they consume oxygen in respiration is converted back into oxygen by plants, algae and bacteria.

Anoxygenic photosynthesis, meanwhile, uses electron donors that are not water and the process does not generate oxygen, according to "Anoxygenic Photosynthetic Bacteria" by LibreTexts . The process typically occurs in bacteria such as green sulfur bacteria and phototrophic purple bacteria. 

The Photosynthesis equation

Though both types of photosynthesis are complex, multistep affairs, the overall process can be neatly summarized as a chemical equation.

The oxygenic photosynthesis equation is: 

6CO2 + 12H2O + Light Energy → C6H12O6 + 6O2 + 6H2O

Here, six molecules of carbon dioxide (CO2) combine with 12 molecules of water (H2O) using light energy. The end result is the formation of a single carbohydrate molecule (C6H12O6, or glucose) along with six molecules each of oxygen and water.

Similarly, the various anoxygenic photosynthesis reactions can be represented as a single generalized formula:

CO2 + 2H2A + Light Energy → [CH2O] + 2A + H2O

The letter A in the equation is a variable, and H2A represents the potential electron donor. For example, "A" may represent sulfur in the electron donor hydrogen sulfide (H2S), according to medical and life sciences news site News Medical Life Sciences . 

How is carbon dioxide and oxygen exchanged?

Plants absorb CO2 from the surrounding air and release water and oxygen via microscopic pores on their leaves called stomata. 

When stomata open, they let in CO2; however, while open, the stomata release oxygen and let water vapor escape. Stomata close to prevent water loss, but that means the plant can no longer gain CO2 for photosynthesis. This tradeoff between CO2 gain and water loss is a particular problem for plants growing in hot, dry environments. 

How do plants absorb sunlight for photosynthesis?

Plants contain special pigments that absorb the light energy needed for photosynthesis.

Chlorophyll is the primary pigment used for photosynthesis and gives plants their green color, according to science education site Nature Education . Chlorophyll absorbs red and blue light and reflects green light. Chlorophyll is a large molecule and takes a lot of resources to make; as such, it breaks down towards the end of the leaf's life, and most of the pigment's nitrogen (one of the building blocks of chlorophyll) is resorbed back into the plant,  When leaves lose their chlorophyll in the fall, other leaf pigments such as carotenoids and anthocyanins begin to show. While carotenoids primarily absorb blue light and reflect yellow, anthocyanins absorb blue-green light and reflect red light, according to Harvard University's The Harvard Forest .

Related: What if humans had photosynthetic skin?

Pigment molecules are associated with proteins, which allow them the flexibility to move toward light and toward one another. A large collection of 100 to 5,000 pigment molecules constitutes an "antenna," according to an article by Wim Vermaas , a professor at Arizona State University. These structures effectively capture light energy from the sun, in the form of photons.

The situation is a little different for bacteria. While cyanobacteria contain chlorophyll, other bacteria, for example, purple bacteria and green sulfur bacteria, contain bacteriochlorophyll to absorb light for anoxygenic photosynthesis, according to " Microbiology for Dummies " (For Dummies, 2019). 

It was previously hypothesized that just a small number of photons would be needed to kickstart photosynthesis, but researchers never successfully observed this first step. However, in 2023, scientists discovered that photosynthesis appears to begin with a single photon. 

The researchers set up an experiment where a photon source spat out two photons at a time. One was absorbed by a detector, while the other hit a bacteria's chloroplast equivalent. When the second photon hit, photosynthesis began. 

After performing the test over 1.5 million times, the researchers confirmed that just one photon is needed to start photosynthesis.

Where in the plant does photosynthesis take place?

Photosynthesis occurs in chloroplasts, a type of plastid (an organelle with a membrane) that contains chlorophyll and is primarily found in plant leaves. 

Chloroplasts are similar to mitochondria , the energy powerhouses of cells, in that they have their own genome, or collection of genes, contained within circular DNA. These genes encode proteins that are essential to the organelle and to photosynthesis.

Inside chloroplasts are plate-shaped structures called thylakoids that are responsible for harvesting photons of light for photosynthesis, according to the biology terminology website Biology Online . The thylakoids are stacked on top of each other in columns known as grana. In between the grana is the stroma — a fluid containing enzymes, molecules and ions, where sugar formation takes place. 

Ultimately, light energy must be transferred to a pigment-protein complex that can convert it to chemical energy, in the form of electrons. In plants, light energy is transferred to chlorophyll pigments. The conversion to chemical energy is accomplished when a chlorophyll pigment expels an electron, which can then move on to an appropriate recipient. 

The pigments and proteins that convert light energy to chemical energy and begin the process of electron transfer are known as reaction centers.

When a photon of light hits the reaction center, a pigment molecule such as chlorophyll releases an electron.

The released electron escapes  through a series of protein complexes linked together, known as an electron transport chain. As it moves through the chain, it generates the energy to produce ATP (adenosine triphosphate, a source of chemical energy for cells) and NADPH — both of which are required in the next stage of photosynthesis in the Calvin cycle. The "electron hole" in the original chlorophyll pigment is filled by taking an electron from water. This splitting of water molecules releases oxygen into the atmosphere.

Light-independent reactions: The Calvin cycle

The Calvin cycle is the three-step process that generates sugars for the plant, and is named after Melvin Calvin , the Nobel Prize -winning scientist who discovered it decades ago. The Calvin cycle uses the ATP and NADPH produced in chlorophyll to generate carbohydrates. It takes plate in the plant stroma, the inner space in chloroplasts.

In the first step of this cycle, called carbon fixation, an enzyme called RuBP carboxylase/oxygenase, also known as rubiso, helps incorporate CO2 into an organic molecule called 3-phosphoglyceric acid (3-PGA). In the process, it breaks off a phosphate group on six ATP molecules to convert them to ADP, releasing energy in the process, according to LibreTexts.

In the second step, 3-PGA is reduced, meaning it takes electrons from six NADPH molecules and produces two glyceraldehyde 3-phosphate (G3P) molecules.

One of these G3P molecules leaves the Calvin cycle to do other things in the plant. The remaining G3P molecules go into the third step, which is regenerating rubisco. In between these steps, the plant produces glucose, or sugar.

Three CO2 molecules are needed to produce six G3P molecules, and it takes six turns around the Calvin cycle to make one molecule of carbohydrate, according to educational website Khan Academy.

There are three main types of photosynthetic pathways: C3, C4 and CAM. They all produce sugars from CO2 using the Calvin cycle, but each pathway is slightly different.

C3 photosynthesis

Most plants use C3 photosynthesis, according to the photosynthesis research project Realizing Increased Photosynthetic Efficiency (RIPE) . C3 plants include cereals (wheat and rice), cotton, potatoes and soybeans. This process is named for the three-carbon compound 3-PGA that it uses during the Calvin cycle. 

C4 photosynthesis

Plants such as maize and sugarcane use C4 photosynthesis. This process uses a four-carbon compound intermediate (called oxaloacetate) which is converted to malate , according to Biology Online. Malate is then transported into the bundle sheath where it breaks down and releases CO2, which is then fixed by rubisco and made into sugars in the Calvin cycle (just like C3 photosynthesis). C4 plants are better adapted to hot, dry environments and can continue to fix carbon even when their stomata are closed (as they have a clever storage solution), according to Biology Online. 

CAM photosynthesis

Crassulacean acid metabolism (CAM) is found in plants adapted to very hot and dry environments, such as cacti and pineapples, according to the Khan Academy. When stomata open to take in CO2, they risk losing water to the external environment. Because of this, plants in very arid and hot environments have adapted. One adaptation is CAM, whereby plants open stomata at night (when temperatures are lower and water loss is less of a risk). According to the Khan Academy, CO2 enters the plants via the stomata and is fixed into oxaloacetate and converted into malate or another organic acid (like in the C4 pathway). The CO2 is then available for light-dependent reactions in the daytime, and stomata close, reducing the risk of water loss. 

Discover more facts about photosynthesis with the educational science website sciencing.com . Explore how leaf structure affects photosynthesis with The University of Arizona . Learn about the different ways photosynthesis can be measured with the educational science website Science & Plants for Schools .  

This article was updated by Live Science managing editor Tia Ghose on Nov. 3, 2022.

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Daisy Dobrijevic joined  Space.com  in February 2022 as a reference writer having previously worked for our sister publication  All About Space  magazine as a staff writer. Before joining us, Daisy completed an editorial internship with the BBC Sky at Night Magazine and worked at the  National Space Centre  in Leicester, U.K., where she enjoyed communicating space science to the public. In 2021, Daisy completed a PhD in plant physiology and also holds a Master's in Environmental Science, she is currently based in Nottingham, U.K.

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• Biology Article
• What is Photosynthesis

What Is Photosynthesis?

“Photosynthesis is the process used by green plants and a few organisms that use sunlight, carbon dioxide and water to prepare their food.”

The process of photosynthesis is used by plants, algae and certain bacteria that convert light energy into chemical energy. The glucose formed during the process of photosynthesis provides two important resources to organisms: energy and fixed carbon.

Read on to explore what is photosynthesis and the processes associated with it.

Site of Photosynthesis

Photosynthesis takes place in special organelles known as chloroplast. This organelle has its own DNA, genes and hence can synthesize its own proteins. Chloroplasts consist of stroma, fluid, and stack of thylakoids known as grana. There are three important pigments present in the chloroplast that absorb light energy, chlorophyll a, chlorophyll b, and carotenoids.

Also Read: Photosynthesis Process

Types of Photosynthesis

There are two different types of photosynthesis:

• Oxygenic photosynthesis
• Anoxygenic photosynthesis

Oxygenic Photosynthesis

Oxygenic photosynthesis is more common in plants, algae and cyanobacteria. During this process, electrons are transferred from water to carbon dioxide by light energy, to produce energy. During this transfer of electrons, carbon dioxide is reduced while water is oxidized, and oxygen is produced along with carbohydrates.

During this process, plants take in carbon dioxide and expel oxygen into the atmosphere.

This process can be represented by the equation:

6CO2+ 12H2O + LIGHT ENERGY → C6H12O6 + 6O2 + 6H2O

Anoxygenic Photosynthesis

This type of photosynthesis is usually seen in certain bacteria, such as green sulphur bacteria and purple bacteria which dwell in various aquatic habitats. Oxygen is not produced during the process.

The anoxygenic photosynthesis can be represented by the equation:

CO2 + 2H2A + LIGHT ENERGY → [CH2O] + 2A + H2O

Also Read:  Difference between Photosynthesis and Respiration

Photosynthesis Apparatus

The photosynthesis apparatus includes the following essential components:

Pigments not only provide colour to the photosynthetic organisms, but are also responsible for trapping sunlight. The important pigments associated with photosynthesis include:

• Chlorophyll: It is a green-coloured pigment that traps blue and red light. Chlorophyll is subdivided into, “chlorophyll a”, “chlorophyll b”, and “chlorophyll c”. “Chlorophyll a” is widely present in all the photosynthetic cells. A bacterial variant of chlorophyll known as bacteriochlorophyll can absorb infrared rays .
• Carotenoids: These are yellow, orange or red-coloured pigments that absorb bluish-green light. Xanthophyll and carotenes are examples of carotenoids.
• Phycobilins: These are present in bacteria and red algae . These are red and blue pigments that absorb wavelength of light that are not properly absorbed by carotenoids and chlorophyll.

Plastids are organelles found in the cytoplasm of eukaryotic photosynthetic organisms. They contain pigments and can also store nutrients. Plastids are of three types:

• Leucoplast: These are colourless, non-pigmented and can store fats and starch.
• Chromoplasts: They contain carotenoids.
• Chloroplasts: These contain chlorophyll and are the site of photosynthesis.

Antennae is the collection of 100 to 5000 pigment molecules that capture light energy from the sun in the form of photons. The light energy is transferred to a pigment-protein complex that converts light energy to chemical energy.

Reaction Centers

The pigment-protein complex responsible for the conversion of light energy to chemical energy forms the reaction centre.

Also Read: Photosynthesis

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Microbe Notes

Photosynthesis: Equation, Steps, Process, Diagram

Photosynthesis is defined as the process, utilized by green plants and photosynthetic bacteria, where electromagnetic radiation is converted into chemical energy and uses light energy to convert carbon dioxide and water into carbohydrates and oxygen.

• The carbohydrates formed from photosynthesis provide not only the necessary energy form the energy transfer within ecosystems, but also the carbon molecules to make a wide array of biomolecules.
• Photosynthesis is a light-driven oxidation-reduction reaction where the energy from the light is used to oxidize water, releasing oxygen gas and hydrogen ions, followed by the transfer of electrons to carbon dioxide, reducing it to organic molecules.
• Photosynthetic organisms are called autotrophs because they can synthesize chemical fuels such as glucose from carbon dioxide and water by utilizing sunlight as an energy source.
• Other organisms that obtain energy from other organisms also ultimately depend on autotrophs for energy.
• One of the essential requirements for photosynthesis is the green pigment ‘chlorophyll’ which is present in the chloroplasts of green plants and some bacteria.
• The pigment is essential for ‘capturing’ sunlight which then drives the overall process of photosynthesis.

Table of Contents

Interesting Science Videos

Photosynthesis equations/reactions/formula

• The process of photosynthesis differs in green plants and sulfur bacteria.
• In plants, water is utilized along with carbon dioxide to release glucose and oxygen molecules.
• In the case of sulfur bacteria, hydrogen sulfide is utilized along with carbon dioxide to release carbohydrates, sulfur, and water molecules.

Oxygenic Photosynthesis

The overall reaction of photosynthesis in plants is as follows:

Carbon dioxide + Water  + solar energy → Glucose + Oxygen

6CO 2 + 6H 2 O  +  solar energy   →   C 6 H 12 O 6 + 6O 2

Carbon dioxide + Water  + solar energy → Glucose + Oxygen + Water

6CO 2 + 12H 2 O+ solar energy    →    C 6 H 12 O 6 + 6O 2 + 6H 2 O

Anoxygenic Photosynthesis

The overall reaction of photosynthesis in sulfur bacteria is as follows:

CO 2 + 2H 2 S + light energy   →    (CH 2 O)  + H 2 O  + 2S

Video Animation: Photosynthesis (Crash Course)

Photosynthetic pigments

• Photosynthetic pigments are the molecules involved in absorbing electromagnetic radiation, transferring the energy of the absorbed photons to the reaction center, resulting in photochemical reactions in the organisms capable of photosynthesis.
• The molecules of photosynthetic pigments are quite ubiquitous and are always composed of chlorophylls and carotenoids.
• In addition to chlorophyll, photosynthetic systems also contain another pigment, pheophytin (bacteriopheophytin in bacteria), which plays a crucial role in the transfer of electrons in photosynthetic systems.
• Moreover, other pigments can be found in particular photosynthetic systems, such as xanthophylls in plants.

Image Source: Simply Science .

Chlorophyll

• Chlorophyll is the pigment molecule, which is the principal photoreceptor in the chloroplasts of most green plants.
• Chlorophylls consist of a porphyrin ring, which is bounded to an ion Mg 2+ , attached to a phytol chain.
• Chlorophylls are very effective photoreceptors because they contain networks of alternating single and double bonds.
• In chlorophyll, the electrons are not localized to a particular atomic nucleus and consequently can more readily absorb light energy.
• In addition, chlorophylls also have solid absorption bands in the visible region of the spectrum.
• Chlorophylls are found either in the cytoplasmic membranes of photosynthetic bacteria, or thylakoid membranes inside plant chloroplasts.

Bacteriorhodopsin

• Bacteriorhodopsin is another class of photosynthetic pigment that exists only in halobacteria.
• It is composed of a protein attached to a retinal prosthetic group.
• This pigment is responsible for the absorption of light photons, leading to a conformational change in the protein, which results in the expulsion of the protons from the cell.

Phycobilins

• Cyanobacteria and red algae employ phycobilins such as phycoerythrobilin and phycocyanobilin as their light-harvesting pigments.
• These open-chain tetrapyrroles have the extended polyene system found in chlorophylls, but not their cyclic structure or central Mg 2+ .
• Phycobilins are covalently linked to specific binding proteins, forming phycobiliproteins, which associate in highly ordered complexes called phycobilisomes that constitute the primary light-harvesting structures in these microorganisms.

Carotenoids

• In addition to chlorophylls, thylakoid membranes contain secondary light-absorbing pigments, or accessory pigments, called carotenoids.
• Carotenoids may be yellow, red, or purple. The most important are β -carotene, which is a red-orange isoprenoid, and the yellow carotenoid lutein.
• The carotenoid pigments absorb light at wavelengths not absorbed by the chlorophylls and thus are supplementary light receptors.

Factors affecting photosynthesis

Blackman formulated the Law of limiting factors while studying the factors affecting the rate of photosynthesis. This Law states that the rate of a physiological process will be limited by the factor which is in the shortest supply. In the same way, the rate of photosynthesis is also affected by a number of factors, which are namely;

• As the intensity of light increases, the rate of light-dependent reactions of photosynthesis and in turn, the rate of photosynthesis increases.
• With increased light intensity, the number of photons falling on a leaf also increases. As a result, more chlorophyll molecules are ionized, and more ATPs and NADH are generated.
• After a point, however, the rate of photosynthesis remains constant as the light intensity increases. At this point, photosynthesis is limited by some other factors.
• Besides, the wavelength of light also affects the rate of photosynthesis.
• Different photosynthetic systems absorb light energy more effectively at different wavelengths.

Carbon dioxide

• An increase in the concentration of carbon dioxide increases the rate at which carbon is incorporated into carbohydrates in the light-independent reactions of photosynthesis.
• Thus, increasing the concentration of carbon dioxide in the atmosphere rapidly increases the rate of photosynthesis up to a point after which it is limited by some other factors.

Temperature

• The light-independent reactions of photosynthesis are affected by changes in temperature as they are catalyzed by enzymes, whereas the light-dependent reactions are not.
• The rate of the reactions increases as the enzymes reach their optimum temperature, after which the rate begins to decrease as the enzymes tend to denature.

Process/ Steps of Photosynthesis

The overall process of photosynthesis can be objectively divided into four steps/ process:

1. Absorption of light

• The first step in photosynthesis is the absorption of light by chlorophylls that are attached to the proteins in the thylakoids of chloroplasts.
• The light energy absorbed is then used to remove electrons from an electron donor like water, forming oxygen.
• The electrons are further transferred to a primary electron acceptor, quinine (Q) which is similar to CoQ in the electron transfer chain.

2. Electron Transfer

• The electrons are now further transferred from the primary electron acceptor through a chain of electron transfer molecules present in the thylakoid membrane to the final electron acceptor, which is usually NADP + .
• As the electrons are transferred through the membrane, protons are pumped out of the membrane, resulting in the proton gradient across the membrane.

3. Generation of ATP

• The movement of protons from the thylakoid lumen to the stroma through the F 0 F 1 complex results in the generation of ATP from ADP and Pi.
• This step is identical to the step of the generation of ATP in the electron transport chain .

4. Carbon Fixation

• The NADP and ATP generated in steps 2 and 3 provide energy, and the electrons drive the process of reducing carbon into six-carbon sugar molecules.
• The first three steps of photosynthesis are directly dependent on light energy and are thus, called light reactions, whereas the reactions in this step are independent of light and thus are termed dark reactions.

Types/ Stages/ Parts of photosynthesis

Figure: Photosynthesis takes place in two stages: light-dependent reactions and the Calvin cycle. Light-dependent reactions, which take place in the thylakoid membrane, use light energy to make ATP and NADPH. The Calvin cycle, which takes place in the stroma, uses energy derived from these compounds to make GA3P from CO 2 . Image Source: OpenStax (Rice University) .

Photosynthesis is divided into two stages based on the utilization of light energy:

1. Light-dependent reactions

• The light-dependent reactions of photosynthesis only take place when the plants/ bacteria are illuminated.
• In the light-dependent reactions, chlorophyll and other pigments of photosynthetic cells absorb light energy and conserve it as ATP and NADPH while simultaneously, evolving O 2 gas.
• In the light-dependent reactions of photosynthesis, the chlorophyll absorbs high energy, short-wavelength light, which excites the electrons present inside the thylakoid membrane.
• The excitation of electrons now initiates the transformation of light energy into chemical energy.
• The light reactions take in two photosystems that are present in the thylakoid of chloroplasts.

Figure: Light-dependent reactions of photosynthesis in the thylakoid membrane of plant cells. Image Source: Wikipedia (Somepics) .

Photosystem II

• Photosystem II is a group of proteins and pigments that work together to absorb light energy and transfer electrons through a chain of molecules until it finally reaches an electron acceptor.
• Photosystem II has a pair of chlorophyll molecules, also known as P680 as the molecules best absorb light of the wavelength 680 nm.
• The P680 donates a pair of electrons after absorbing light energy, resulting in an oxidized form of P680.
• Finally, an enzyme catalyzes the splitting of a water molecule into two electrons, two hydrogen ion, and oxygen molecules.
• This pair of electrons then are transferred to P680, causing it to return to its initial stage.

Photosystem I

• Photosystem I is a similar complex like photosystem II except for that photosystem I have a pair of chlorophyll molecules known as P700 as they best absorb the wavelength of 700 nm.
• As photosystem I absorb light energy, it also becomes excited and transfers electrons.
• The now oxidized form of P700 then accepts an electron from photosystem II, restring back to its initial stage.
• The electrons from photosystem I are then passed in a series of redox reactions through the protein ferredoxin.
• The electrons finally reach NADP + , reducing them to NADPH.

2 H 2 O + 2 NADP +  + 3 ADP + 3 P i  + light → 2 NADPH + 2 H +  + 3 ATP + O 2

Video Animation: The Light Reactions of Photosynthesis (Ricochet Science)

2. Light independent reactions (Calvin cycle)

Light independent reactions of photosynthesis are anabolic reactions that lead to the formation of a sex-carbon compound, glucose in plants. The reactions in this stage are also termed dark reactions as they are not directly dependent on the light energy but do require the products formed from the light reactions.

Figure: Overview of the Calvin cycle pathway. Image Source: Wikipedia (Mike Jones) .

This stage consists of 3 further steps that lead to carbon fixation/ assimilation.

Step 1: Fixation of CO 2 into 3-phosphoglycerate

• In this step, one CO 2 molecule is covalently attached to the five-carbon compound ribulose 1,5-biphosphate catalyzed by the enzyme ribulose 1,5-biphosphate carboxylase, also called rubisco.
• The attachment results in the formation of an unstable six-carbon compound that is then cleaved to form two molecules of 3-phosphoglycerate.

Step 2: Conversion of 3-phosphoglycerate to glyceraldehydes 3-phosphate

• The 3-phosphoglycerate formed in step 1 is converted to glyceraldehyde 3-phosphate by two separate reactions.
• At first, enzyme 3-phosphoglycerate kinase present in the stroma catalyzes the transfer of a phosphoryl group from ATP to 3-phosphoglycerate, yielding 1,3-bisphosphoglycerate.
• Next, NADPH donates electrons in a reaction catalyzed by the chloroplast-specific isozyme of glyceraldehyde 3-phosphate dehydrogenase, producing glyceraldehyde 3-phosphate and phosphate (Pi).
• Most of the glyceraldehyde 3-phosphate thus produced is used to regenerate ribulose 1,5-bisphosphate.
• The rest of the glyceraldehyde is either converted to starch in the chloroplast and stored for later use or is exported to the cytosol and converted to sucrose for transport to growing regions of the plant.

Step 3: Regeneration of ribulose 1,5-biphosphate from triose phosphates

• The three-carbon compounds formed in the previous steps are then converted into the five-carbon compound, ribulose 1,5-biphosphate through a series of transformations with intermediates of three-, four,-, five-, six-, and seven-carbon sugar.
• As the first molecules in the process, if regenerated, this stage of photosynthesis results in a cycle (Calvin cycle).

3 CO 2 + 9 ATP + 6 NADPH + 6 H +     →     glyceraldehyde-3-phosphate (G3P) + 9 ADP + 8 P i  + 6 NADP +  + 3 H 2 O

A G3P molecule contains three fixed carbon atoms, so it takes two G3Ps to build a six-carbon glucose molecule. It would take six turns of the cycle to produce one molecule of glucose.

Video Animation: The Calvin Cycle (Ricochet Science)

Products of Photosynthesis

The outcomes of light-dependent reactions of photosynthesis are:

The products of light-independent reactions (Calvin cycle) of photosynthesis are:

• glyceraldehyde-3-phosphate (G3P) / Glucose (carbohydrates)

The overall products of photosynthesis are:

• Glucose (carbohydrates)
• Sulfur (in photosynthetic sulfur bacteria)

Photosynthesis Examples

Photosynthesis in green plants or oxygenic bacteria.

• In plants and oxygenic bacteria like cyanobacteria, photosynthesis takes place in the presence of green pigment, chlorophyll.
• It takes place in the thylakoids of the chloroplasts, resulting in products like oxygen gas, glucose, and water molecules.
• Most of the glucose units in plants are linked to form starch or fructose or even sucrose.

Photosynthesis in sulfur bacteria

• In purple sulfur bacteria, photosynthesis takes place in the presence of hydrogen sulfur rather than water.
• Some of these bacteria like green sulfur bacteria have chlorophyll whereas other purple sulfur bacteria have carotenoids as photosynthetic pigments.
• The result of photosynthesis in these bacteria are carbohydrates (not necessarily glucose), sulfur gas, and water molecules.

Importance of photosynthesis

• Photosynthesis is the primary source of energy in autotrophs where they make their food by utilizing carbon dioxide, sunlight, and photosynthetic pigments.
• Photosynthesis is equally essential for heterotrophs, as they derive their energy from the autotrophs.
• Photosynthesis in plants is necessary to maintain the oxygen levels in the atmosphere.
• Besides, the products of photosynthesis contribute to the carbon cycle occurring in the oceans, land, plants, and animals.
• Similarly, it also helps maintain a symbiotic relationship between plants, animals, and humans.
• Sunlight or solar energy is the primary source of all other forms of energy on earth, which is utilized through the process of photosynthesis.

Artificial photosynthesis

Artificial photosynthesis is a chemical process that mimics the biological process of utilization of sunlight, water and carbon dioxide to produce oxygen and carbohydrates.

Image Source: Phys .

• In artificial photosynthesis, photocatalysts are utilized that are capable of replicating the oxidation-reduction reactions taking place during natural photosynthesis.
• The main function of artificial photosynthesis is to produce solar fuel from sunlight that can be stored and used under conditions, where sunlight is not available.
• As solar fuels are prepared, artificial photosynthesis can be used to produce just oxygen from water and sunlight, resulting in clean energy production.
• The most important part of artificial photosynthesis is the photocatalytic splitting of a water molecule, resulting in oxygen and large quantities of hydrogen gas.
• Further, light-driven carbon reduction can also be performed to replicate the process of natural carbon fixation, resulting in carbohydrates molecules.
• Thus, artificial photosynthesis has applications in the production of solar fuels, photoelectrochemistry, engineering of enzymes, and photoautotrophic microorganisms for the production of microbial biofuel and biohydrogen from sunlight.

Video Animation: Learning from leaves: Going green with artificial photosynthesis

Photosynthesis vs. Cellular respiration

Image Source: Khan Academy .

Video Animation: Photosynthesis vs. Cellular Respiration Comparison (BOGObiology)

FAQs (Revision Questions)

Where does photosynthesis occur? Photosynthesis occurs in the thylakoid membrane of the chloroplasts.

What are the products of photosynthesis? The products of photosynthesis are carbohydrates (glucose), oxygen, and water molecules.

What are the reactants of photosynthesis? The reactants of photosynthesis are carbon dioxide, water, photosynthetic pigments, and sunlight.

How are photosynthesis and cellular respiration related? Photosynthesis and cellular respiration are essentially the reverses of one another where photosynthesis is an anabolic process resulting in the formation of organic molecules. In contrast, cellular respiration is a catabolic process resulting in the breaking down of organic molecules to release energy.

• Berg JM, Tymoczko JL, Stryer L. Biochemistry. 5th edition. New York: W H Freeman; 2002. Section 17.2, Entry to the Citric Acid Cycle and Metabolism Through It Are Controlled.Available from: https://www.ncbi.nlm.nih.gov/books/NBK22347/
• Nelson DL and Cox MM. Lehninger Principles of Biochemistry. Fourth Edition.
• Montero F. (2011) Photosynthetic Pigments. In: Gargaud M. et al. (eds) Encyclopedia of Astrobiology. Springer, Berlin, Heidelberg
• Lodish H, Berk A, Zipursky SL, et al. Molecular Cell Biology. 4th edition. New York: W. H. Freeman; 2000. Section 16.3, Photosynthetic Stages and Light-Absorbing Pigments.Available from: https://www.ncbi.nlm.nih.gov/books/NBK21598/

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Photosynthesis

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Photosynthesis is a vital biological process through which green plants, algae, and certain bacteria convert light energy into chemical energy. Using sunlight, these organisms transform carbon dioxide and water into glucose and oxygen, substances crucial for their growth and the sustenance of life on Earth. This process not only fuels the organisms themselves but also supports life on the planet by providing oxygen and forming the base of the food chain. Understanding photosynthesis is essential for comprehending how life thrives on Earth, influencing fields ranging from agriculture to energy production.

Definition of Photosynthesis

Photosynthesis is a fundamental biological process through which green plants, algae, and certain bacteria convert light energy into chemical energy. This transformation occurs primarily in the chloroplasts of plant cells where chlorophyll, the pigment responsible for the green color of plants, captures sunlight. The captured light energy is then used to synthesize glucose from carbon dioxide (CO2) and water (H2O), releasing oxygen (O2) as a byproduct. This process not only fuels the plant’s own cellular activities but also provides the base of the food chain for other organisms.

Where Does Photosynthesis Occur?

Photosynthesis primarily occurs in the leaves of plants, although it can also take place in any parts of a plant that contain green pigments, typically in the stems and young branches. The leaves are the main site of photosynthesis due to their structure and accessibility to sunlight.

Photosynthesis Process

Photosynthesis stands as a crucial biological process through which plants, algae, and certain bacteria convert sunlight into chemical energy, fueling their activities and growth. This process not only supports the organisms performing it but also sustains life on Earth by producing oxygen and forming the basis of the food chain.

Key Stages of Photosynthesis

Photosynthesis occurs primarily in two stages: the light-dependent reactions and the light-independent reactions (Calvin cycle).

Light-Dependent Reactions

These reactions take place in the thylakoid membranes of the chloroplasts. Here, sunlight drives the process, initiating the flow of electrons through a series of proteins known as the electron transport chain. Plants absorb sunlight using pigment molecules, with chlorophyll being the most prominent. This absorption energizes electrons, which the chloroplasts then use to convert water (H₂O) into oxygen (O₂). As a result, this stage releases oxygen as a byproduct and generates ATP and NADPH, energy carriers that the next stage of photosynthesis uses.

Location: Thylakoid Membranes

Within the chloroplasts, thylakoid membranes house the light-dependent reactions. These membranes are rich in pigments like chlorophyll that capture light energy, crucial for water photolysis and energy molecule production.

Steps in Light-Dependent Reactions

• Photon Absorption : When photons strike the chlorophyll in the photosystem II (PSII) complex, embedded in the thylakoid membrane, they excite electrons to a higher energy state.
• Water Splitting (Photolysis) : To replace these excited electrons, enzymes split water molecules into oxygen, protons, and electrons. The process releases oxygen as a byproduct.
• Electron Transport Chain (ETC) : The electrons travel down an electron transport chain, which comprises a series of proteins within the thylakoid membrane. The movement of electrons through the chain helps pump protons from the stroma into the thylakoid space, thereby creating a proton gradient.
• ATP Formation : This proton gradient enables ATP synthase to synthesize ATP from ADP and inorganic phosphate through chemiosmosis, as it uses the flow of protons back into the stroma.
• Photosystem I (PSI) : Upon reaching PSI, the electrons get re-energized by more absorbed light. These re-energized electrons then reduce NADP+ to NADPH, which serves as a critical electron and hydrogen carrier in the Calvin cycle.
• Output of Light-Dependent Reactions : ATP and NADPH, produced during these reactions, enter the Calvin cycle to assist in carbon dioxide fixation into glucose. Meanwhile, the oxygen produced during water splitting exits as a waste product.

Significance of Light-Dependent Reactions

The light-dependent reactions are vital as they provide the necessary energy carriers (ATP and NADPH) for the Calvin cycle. They also maintain the oxygen level in the atmosphere, which is critical for the survival of aerobic life on Earth.

Calvin Cycle (Light-Independent Reactions)

The Calvin cycle unfolds in the stroma, the fluid-filled space surrounding the thylakoid membranes. It does not require light directly. Instead, it uses the ATP and NADPH from the light-dependent reactions to convert carbon dioxide (CO₂) from the air into organic molecules. During this cycle, the enzyme RuBisCO incorporates CO₂ into an organic molecule, starting a series of chemical reactions that regenerate the starting molecule with the production of glucose and other carbohydrates.

Key Steps of the Calvin Cycle

• Carbon Fixation : The cycle begins when the enzyme RuBisCO incorporates carbon dioxide into a five-carbon molecule, ribulose bisphosphate (RuBP). This reaction produces a six-carbon compound that immediately splits into two three-carbon molecules of 3-phosphoglycerate (3-PGA).
• Reduction Phase : Each molecule of 3-PGA then receives a phosphate group from ATP, becoming 1,3-bisphosphoglycerate. Next, NADPH donates electrons to these molecules, reducing them to glyceraldehyde-3-phosphate (G3P), a sugar. This step consumes the ATP and NADPH generated during the light-dependent reactions.
• Release of One Glucose Molecule : For every six molecules of CO2 that enter the cycle, twelve molecules of G3P form. However, only two G3P molecules leave the cycle to contribute towards forming glucose, while the remaining ten G3P molecules proceed to the next step.
• Regeneration of RuBP : The ten remaining G3P molecules undergo a series of transformations that require further ATP. These transformations regenerate RuBP, enabling the cycle to process new carbon dioxide molecules continuously.

Significance of the Calvin Cycle

The Calvin Cycle is crucial for synthesizing glucose, which plants use as an energy source to fuel various cellular activities and growth. The glucose also serves as a building block for other essential biomolecules such as cellulose and starch. Additionally, this cycle plays a pivotal role in the global carbon cycle, as it is the primary pathway through which atmospheric carbon dioxide transforms into organic compounds within plants.

Importance of Photosynthesis

Photosynthesis is vital for life on Earth. It provides the primary energy source for all ecosystems, where plants form the base of the food web and create biomass from inorganic substances. Moreover, photosynthesis is responsible for the oxygen that makes up a significant portion of the Earth’s atmosphere and supports aerobic life forms.

Photosynthesis Equation

The formula for photosynthesis is central to understanding how plants convert solar energy into chemical energy. Here is the equation:

6𝐶𝑂2+6𝐻2𝑂+𝑙𝑖𝑔ℎ𝑡𝑒𝑛𝑒𝑟𝑔𝑦→𝐶6𝐻12𝑂6+6𝑂26 CO 2​+6 H 2​ O + lightenergy → C 6​ H 12​ O 6​+6 O 2​

Explanation of the Photosynthesis Equation

This equation represents the overall process by which plants, algae, and certain bacteria produce glucose and oxygen from carbon dioxide and water, using energy from sunlight. Here’s a breakdown of each component:

Carbon Dioxide (6CO2)

Tiny pores in the leaves called stomata take carbon dioxide from the air. Carbon dioxide is one of the key reactants in the process.

Water (6H2O)

The roots absorb water and transport it to the leaves, providing the source of electrons and protons necessary for the chemical reactions of photosynthesis. Water molecules split to produce oxygen.

Light Energy

Chlorophyll and other pigments in the chloroplasts capture light energy, converting it into chemical energy in the form of ATP and NADPH. These energy carriers then power the later stages of photosynthesis.

Glucose (C6H12O6)

The plant uses the sugar produced by photosynthesis as an energy source. It can use this sugar immediately, store it, or convert it into other necessary substances for growth.

Oxygen (6O2)

Oxygen releases through the stomata as a byproduct, playing a critical role in the Earth’s atmosphere and supporting the survival of aerobic organisms.

Causes of Photosynthesis

Light energy, especially from the sun, triggers photosynthesis. Pigments in the plant, mainly chlorophyll, absorb mostly blue and red wavelengths of light, crucial for converting ADP and inorganic phosphate into ATP, and NADP+ into NADPH, which are used in glucose synthesis.

Chlorophyll and Other Pigments

Chlorophyll primarily absorbs light. Other pigments like carotenoids and phycobilins help capture energy from sunlight, absorbing light wavelengths that chlorophyll cannot. These pigments are critical in harnessing the light energy required to drive the reactions of photosynthesis.

Water (H2O)

Water acts as an electron donor in the light-dependent reactions of photosynthesis. The process of photolysis splits water molecules, releasing electrons, hydrogen ions, and oxygen. The reactions use the electrons and hydrogen ions to produce glucose, while plants release oxygen as a byproduct.

Carbon Dioxide (CO2)

The stomata in the leaves absorb CO2 from the atmosphere. It is a critical substrate in the Calvin cycle (light-independent reactions), where it fixes into glucose using the ATP and NADPH produced in the light-dependent reactions.

Various enzymes facilitate the chemical reactions involved in photosynthesis. For example, the enzyme Rubisco plays a pivotal role in fixing carbon dioxide during the Calvin cycle, converting it into glucose. Enzymes ensure that the photosynthetic reactions occur efficiently and at a sufficient rate to meet the plant’s needs.

Cellular Structures (Chloroplasts)

Chloroplasts, specialized organelles in plant and algal cells, house the biochemical machinery necessary for photosynthesis. The thylakoid membranes in chloroplasts provide a framework for light-dependent reactions, while the surrounding stroma hosts the Calvin cycle.

Temperature and pH

Temperature and pH levels also influence the rate of photosynthesis. The enzymatic reactions involved are sensitive to temperature and have optimal pH ranges. Deviations from these optimal conditions can slow down or inhibit the process.

What Cells and Organelles Are Involved in Photosynthesis?

Photosynthesis, a critical process through which green plants, algae, and some bacteria convert light energy into chemical energy, occurs largely in specialized cells and organelles designed to maximize the efficiency of light capture and conversion. The primary cells and organelles involved in photosynthesis are mesophyll cells, chloroplasts, and, more specifically, structures within the chloroplasts including the thylakoid membranes and stroma.

Mesophyll Cells

Mesophyll cells in plant leaves primarily conduct photosynthesis. These cells contain high concentrations of chloroplasts, the essential organelles where photosynthesis takes place. There are two types of mesophyll cells:

Palisade Mesophyll

These cells, located directly under the leaf surface, are elongated and densely packed with chloroplasts, primarily absorbing light and conducting photosynthesis.

Spongy Mesophyll

These cells, found below the palisade mesophyll, aid in gas exchange and also contain chloroplasts contributing to photosynthesis.

Chloroplasts

Chloroplasts are the key organelles where photosynthesis occurs. These double-membraned structures contain their own DNA and can replicate independently within the cell. Inside chloroplasts, two major stages of photosynthesis—the light-dependent reactions and the Calvin cycle—take place in different components:

Thylakoid Membranes

These membrane-bound structures, stacked into grana within chloroplasts, contain chlorophyll, essential for absorbing sunlight. The light-dependent reactions of photosynthesis occur here, converting sunlight into chemical energy in the form of ATP and NADPH.

This fluid-filled space surrounds the thylakoid membranes inside the chloroplast. Here, the Calvin cycle, also known as the light-independent reactions, occurs. It uses the ATP and NADPH produced by the light-dependent reactions to convert carbon dioxide into glucose, serving as an energy storage molecule for the plant.

Importance of Each Component

Each part within the mesophyll cells and chloroplasts plays a crucial role in the process of photosynthesis:

• Mesophyll cells ensure that chloroplasts are in the optimal position to receive sunlight and facilitate gas exchange, which is vital for photosynthesis.
• Chloroplasts function as the site of the photosynthesis process, housing all the necessary molecular machinery.
• Thylakoid membranes are critical for capturing light and transforming it into usable energy.
• Stroma provides the enzymatic playground for synthesizing organic molecules from carbon dioxide and water.

Factors Influencing the Rate of Photosynthesis

1. light intensity.

• Impact : The rate of photosynthesis typically increases as light intensity rises, up to a certain point. Beyond this point, the process plateaus as other factors become limiting.
• Explanation : Light provides the energy needed for photosynthesis. More light equates to more energy available to drive the chemical reactions involved.

2. Carbon Dioxide Concentration

• Impact : Increasing the concentration of carbon dioxide can enhance the rate of photosynthesis, until the process is constrained by another factor.
• Explanation : Carbon dioxide is a raw material used in the formation of glucose during photosynthesis. Higher concentrations can increase the rate of carbon fixation in the Calvin cycle.

3. Temperature

• Impact : Photosynthesis is temperature-dependent, with the rate increasing up to an optimal temperature and then rapidly decreasing at higher temperatures.
• Explanation : Enzymatic reactions that drive photosynthesis perform optimally within a certain temperature range. Too high or too low temperatures can denature these enzymes, reducing the efficiency of photosynthesis.

4. Water Availability

• Impact : Water stress can severely limit the rate of photosynthesis.
• Explanation : Water is not only a reactant in the chemical equation of photosynthesis but also essential for the plant’s overall health and turgidity. Lack of water can lead to stomatal closure to conserve water, thereby reducing CO2 uptake.

5. Quality of Light

• Impact : Different wavelengths of light affect photosynthesis differently. Blue and red lights are most effective in driving photosynthesis.
• Explanation : Chlorophyll, the primary pigment in photosynthesis, absorbs blue and red light more efficiently than other wavelengths.

6. Chlorophyll Content

• Impact : The amount of chlorophyll in leaves affects their ability to capture light energy.
• Explanation : More chlorophyll molecules increase the capacity for light absorption, enhancing the photosynthetic rate.

7. Leaf Anatomy and Orientation

• Impact : Leaf structure and positioning can influence light capture and gas exchange, impacting photosynthesis.
• Explanation : Leaves arranged to maximize light capture and minimize overlap can more effectively convert light energy into chemical energy.

Energy Efficiency of Photosynthesis

Measuring energy efficiency.

The energy efficiency of photosynthesis generally refers to the percentage of solar energy that plants convert into the chemical energy of sugars. Solar energy strikes the Earth with a power of about 1000 watts per square meter at noon on a clear day. Plants absorb only a fraction of this energy, primarily using the visible light spectrum.

Factors Affecting Efficiency

Several factors impact the energy efficiency of photosynthesis:

• Pigment Absorption : Chlorophyll, the primary pigment in plants, absorbs blue and red light effectively but reflects green light, which is why plants appear green. This selective absorption limits the range of light energy plants can use.
• Photosynthetic Active Radiation : Only about 45% of the sunlight’s energy is in the form of photosynthetically active radiation (PAR). Plants primarily use this portion for photosynthesis.
• Energy Conversion Process : The complex series of reactions in photosynthesis includes losses due to reflection, respiration, and heat production.

Calculating Efficiency

• Theoretical Maximum Efficiency : Research suggests that the theoretical maximum efficiency of photosynthesis in converting solar energy into biomass is around 11% under ideal conditions. This accounts for the energy absorbed and utilized in the formation of glucose.
• Typical Real-World Efficiency : In real-world conditions, the efficiency is much lower. On average, photosynthesis converts only about 0.1% to 2% of solar energy into biomass. This range varies significantly with the type of plant, environmental conditions, and time of year.

Implications of Efficiency

Despite its relatively low energy efficiency, photosynthesis is incredibly effective in supporting life on Earth:

• Global Scale : Annually, photosynthesis captures approximately 130 terawatts of energy as biomass, more than six times the current power consumption of human civilization.
• Ecosystem and Climate : The process is essential for carbon capture, which helps regulate atmospheric CO₂ levels and mitigate climate change.
• Agricultural Productivity : Understanding and improving the efficiency of photosynthesis could lead to increased crop yields and better food security globally.

Enhancing Photosynthetic Efficiency

Scientists are researching ways to enhance the efficiency of photosynthesis to benefit food production and bioenergy. These efforts include genetically modifying plants to absorb light more effectively, bypassing inefficient steps in natural photosynthesis, and developing artificial photosynthesis systems that could one day surpass the efficiency of natural photosynthesis.

C4 Photosynthesis

C4 photosynthesis is a highly efficient photosynthetic pathway that some plants use to overcome the limitations of the standard C3 pathway, especially under conditions of drought, high temperatures, and limited nitrogen or CO2. In C4 photosynthesis, plants capture CO2 in the mesophyll cells and then transport it to the bundle-sheath cells where the Calvin cycle occurs. This process minimizes photorespiration, an energy-wasting process that occurs in C3 plants. Key enzymes like PEP carboxylase initially fix CO2 into a four-carbon compound, which is why we call it C4 photosynthesis. This adaptation allows C4 plants, such as maize and sugarcane, to photosynthesize more efficiently under extreme conditions compared to C3 plants.

What is Photosynthesis in Short Answer?

Photosynthesis is the process where plants use sunlight to produce energy from water and carbon dioxide, releasing oxygen.

How Do You Explain Photosynthesis?

Photosynthesis transforms sunlight into chemical energy, enabling plants to create glucose and oxygen from water and CO2.

What is the Main Process of Photosynthesis?

The main process of photosynthesis involves converting light energy into chemical energy, which plants use to make glucose and release oxygen.

How to Start Photosynthesis?

Photosynthesis begins when chlorophyll in plant cells absorbs sunlight, initiating energy conversion that powers chemical reactions.

What is the Simplest Definition of Photosynthesis?

Photosynthesis is the process by which plants make their own food using sunlight, water, and carbon dioxide.

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Introduction

Photosynthesis is a process by which plants, algae, and certain microorganisms transform light energy from the sun into the chemical energy of food. During photosynthesis, energy from sunlight is harnessed and used to convert carbon dioxide and water into organic compounds—namely sugar molecules—and oxygen. The process enables photosynthetic organisms to change light energy into a form of energy—the chemical energy in sugars—that their cells can store and use to grow and thrive. This chemical energy is transferred to animals that eat plants as well as other photosynthetic organisms, and to the animals that eat other animals ( see food chain ). The oxygen produced through photosynthesis is released into the atmosphere (in the case of aquatic organisms, into the aquatic environment) and used by organisms for cellular respiration .

The importance of photosynthesis to life on Earth cannot be overstated. Photosynthesis directly and indirectly supplies the food we eat and the air we breathe. Without it, the Earth’s fundamental food supply would not be replenished. With less food available, most organisms would disappear, and the planet would become devoid of oxygen. ( See also ecosystem .)

An overview of photosynthesis is presented in the section that follows. For a detailed review of the chemical reactions in photosynthesis, see Photosynthesis in Detail in this article.

Overview of Photosynthesis

Photosynthesis is a chemical process in which light energy from the sun drives a series of chemical reactions between carbon dioxide and water, forming glucose (a simple sugar) and oxygen as end products. The overall process can be expressed as follows:

carbon dioxide + water + light energy → glucose + oxygen

The reaction requires chlorophyll, a green pigment found in all organisms that can undergo photosynthesis. Chlorophyll plays a critical role in capturing energy from incoming sunlight and transferring it to the chemical compounds involved in photosynthesis. The chlorophyll in plant cells is contained in organelles called chloroplasts. All photosynthesis in plants takes place within the chloroplasts, which are concentrated in the palisade cells of leaves.

Photosynthesis proceeds through a series of steps. Water enters the plant through the roots and travels up to the leaves. Carbon dioxide enters the leaves through tiny pores called stomata ( see plant ). Inside the chloroplasts, chlorophyll captures energy from sunlight; this energy then drives a series of chemical reactions between carbon dioxide and water, changing them into glucose and oxygen.

The oxygen produced in photosynthesis is released into the air as a waste product. The plant may burn some of the glucose immediately for energy to power cellular activities ( see cellular respiration ). Some glucose molecules may be converted to other sugars, such as fructose or sucrose, or linked together to form cellulose —a large complex carbohydrate needed to build and repair cell walls. Most of the glucose produced in photosynthesis is linked together to form starch , a large complex carbohydrate. Plants store starch in their tissues and break it down into glucose molecules when they need energy.

Photosynthesis in Detail

In photosynthesis, light energy reacts with six molecules of carbon dioxide (CO 2 ) and six molecules of water (H 2 O) to produce one molecule of glucose (C 6 H 12 O 6 ) and six molecules of oxygen (O 2 ). The chemical equation for photosynthesis is as follows:

6CO 2  + 6H 2 O +  light  →  C 6 H 12 O 6  +  6O 2

The actual process, however, is far more complex than the equation might suggest. The equation indicates that atmospheric carbon dioxide, CO 2 , is “fixed,” or converted from a gas to the solid sugar glucose (C 6 H 12 O 6 ), water (H 2 O) is consumed, and oxygen (O 2 ) is liberated. Yet this seemingly simple reaction consists of two distinctly different processes. The first is photochemical and the second is biochemical; they are the so-called light and dark, or light-independent, reactions. The rate of these reactions is controlled by various regulatory enzymes as well as by environmental conditions such as light intensity, temperature, and the availability of carbon dioxide, water, and certain minerals.

Light Reaction

All photosynthetic organisms—with the exception of a small group of bacteria—contain various forms of the green pigment chlorophyll, which is critical in the transfer of energy from light to chemical compounds. Other pigments may also be involved, depending on the type of organism undergoing photosynthesis. Light for photosynthesis is harvested from the visible spectrum ( see color ; light ), and different pigments absorb the light from a particular range within that spectrum. The result is a form of light-absorption array, or antenna, that neatly matches the entire visible spectrum.

During the light reaction, incoming light energy is absorbed, or captured, by chlorophyll pigments in the cell. Each photon, or “particle” of light, then undergoes a process called free-charge separation. In this process an electron (e – ) is separated from the chlorophyll molecule and is passed, at a higher energy, to a carrier molecule, thereby converting the energy of the photon into chemical energy. The electrons lost by the chlorophyll molecules are replaced by electrons that split off from water. This process, called photolysis, forms oxygen as a waste product. The photolytic reaction can be described as follows:

H 2 O → 2H + + 2e – + 1 / 2 O 2

Two such free-charge separations, called photoacts, are connected in series. As electrons pass between photoacts, the energy-rich compound adenosine triphosphate (ATP) is formed by the addition of an inorganic phosphate group (P i ) to a molecule of adenosine diphosphate (ADP), and the electron loses energy. This process is called photophosphorylation, and it can be described as follows:

ADP + P i  → ATP + H 2 O

In the second photoact, the electron-acceptor compound nicotinamide adenine dinucleotide phosphate (NADP + ) is reduced—that is, it gains electrons—to form the electron donor compound NADPH:

NADP +  + H +  + 2e –  → NADPH

The compounds ATP and NADPH are used in the next stage of photosynthesis, the dark, or light-independent, reaction. In nature, for every ten photons absorbed, two to three molecules of ATP and two molecules of NADPH are formed. This translates into an energy conversion efficiency of about 38 percent.

Dark (Light-Independent) Reaction

In the dark reaction of photosynthesis, the ATP and NADPH formed in the light reaction are used to transform inorganic carbon dioxide (CO 2 ) into organic carbon compounds, a process called carbon fixation. The term dark reaction is misleading because the process can take place in both light and darkness. For that reason, many people refer to this process as “light independent” rather than “dark.”

Carbon fixation takes place as a series of steps in a complex biochemical cycle known as the Calvin-Benson cycle. This process produces intermediate sugar phosphate compounds that are then used to synthesize glucose. The process begins with three molecules of carbon dioxide (CO 2 ) and produces a net output of one molecule of a 3-carbon sugar phosphate called glyceraldehyde-3-phosphate (Gal3P). The process requires energy from nine molecules of ATP and electrons from six molecules of NADPH. The overall process can be summarized as follows:

3 CO 2 + 9 ATP + 6 NADPH → Gal3P + 6 NADP + + 9 ADP + 8 P i

The cycle begins with the reaction of the 5-carbon sugar ribulose bisphosphate (RuBP) with carbon dioxide (CO2), producing the intermediate 3-carbon molecule phosphoglycerate (PGA). The reaction is brought about, or catalyzed, by the enzyme ribulose bisphosphate carboxylase/oxygenase (RuBisCO):

3 CO 2 + 3 RuBP → 6 PGA

Each PGA molecule undergoes phosphorylation, gaining a phosphate group from ATP, and is reduced with NADPH, thereby gaining an electron. These reactions ultimately produce six molecules of glyceraldehyde-3-phosphate:

6 PGA + 6 ATP + 6 NADPH → 6 Gal3P

Five molecules of Gal3P are converted to form three molecules of the 5-carbon RuBP. The process requires energy from 3 ATP:

5 Gal3P + 3 ATP → 3 RuBP + 3 ADP

The RuBP molecules reenter the cycle, ready to react with new molecules of carbon dioxide.

The remaining molecule of the 3-carbon Gal3P can combine with another to form the 6-carbon sugar glucose:

2 Gal3P → C 6 H 12 O 6

The glucose can then be converted to other sugars, such as fructose or sucrose, or it can be linked to form complex carbohydrates, such as starch or cellulose. Glucose can also be used in other synthetic reactions to form fats or proteins as needed by the plant. ( See also carbohydrate ; plant .)

Begun in the 19th century, the investigation of the mechanism of photosynthesis is still continuing today. A continuing area of research since the 1970s has involved the use of genetic engineering to alter specific photosynthetic mechanisms and improve crop productivity. However, the complex factors in agriculture and in the genetic manipulation of crops have frustrated scientists; by the early 21st century, genetic engineering had not yet been shown to improve crop yields.

An interesting area of research has been the discovery that certain animals are capable of converting light energy into chemical energy on a small scale. For example, the emerald green sea slug (Elysis chlorotica) acquires chloroplasts from the algae it eats. When the slug accumulates enough chloroplasts, it can undergo photosynthesis and no longer needs to ingest food.

Joseph S. Ramus

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photosynthesis

Definition of photosynthesis

Did you know.

Photosynthesis Has Greek Roots

The Greek roots of photosynthesis combine to produce the basic meaning "to put together with the help of light". Photosynthesis is what first produced oxygen in the atmosphere billions of years ago, and it's still what keeps it there. Sunlight splits the water molecules (made of hydrogen and oxygen) held in a plant's leaves and releases the oxygen in them into the air. The leftover hydrogen combines with carbon dioxide to produce carbohydrates, which the plant uses as food—as do any animals or humans who might eat the plant.

Examples of photosynthesis in a Sentence

These examples are programmatically compiled from various online sources to illustrate current usage of the word 'photosynthesis.' Any opinions expressed in the examples do not represent those of Merriam-Webster or its editors. Send us feedback about these examples.

Word History

1898, in the meaning defined above

Dictionary Entries Near photosynthesis

photosynthate

photosynthetic ratio

Cite this Entry

“Photosynthesis.” Merriam-Webster.com Dictionary , Merriam-Webster, https://www.merriam-webster.com/dictionary/photosynthesis. Accessed 29 Jun. 2024.

Kids Definition

Kids definition of photosynthesis, medical definition, medical definition of photosynthesis, more from merriam-webster on photosynthesis.

Nglish: Translation of photosynthesis for Spanish Speakers

Britannica.com: Encyclopedia article about photosynthesis

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Photosynthesis

Photosynthesis for kids - learn all about photosynthesis, what is photosynthesis.

Photosynthesis = “Photo” + “Synthesis”. “Photo” which means light, “Synthesis” which means putting together.

Just think, plants cannot move, but they need food to survive and grow. But how do they manage their food or who helps them?

The answer is, plants are the most independent living thing on earth. They make their own food without any help from people.

Plants use sunlight, water, CO2 ( carbon dioxide i.e. present in air) to make their food. And this process of making food is called. photosynthesis . Now we will explore more about photosynthesis for kids.

✍ How Photosynthesis Works Step by Step?

Let’s explore how photosynthesis works step by step. For photosynthesis, plants need 3 essential things.  Water and Minerals CO2 (Carbon Dioxide) Sunlight Step 1 -The root of the plant takes water and minerals from the soil . There are veins in the plant named Xylem , transport water and minerals to the whole plant, including its leaves.

The leaves are the food factory of the plant. Because the photosynthesis process takes place in the leaves of plants. So, every necessary thing needs to reach the leaves to make food .

Step 2 – Carbon dioxide (CO2) from the air enters through tiny pores (holes) in the leaves. These pores are called stomata.

We can say that plants also breathe like us through stomata. The only difference is, we breathe in oxygen and breathe out carbon dioxide but plants breathe in carbon dioxide and breathe out oxygen. What a fantastic understanding between us and plants!!

Step 3 – The leaves are basically made of tiny cells . If we see the structure of Plant Cells , we will find that inside these cells there is a small part called chloroplasts . Each chloroplast contains a green substance called Chlorophyll , which actually absorbs energy from sunlight.

Did you know? Chlorophyll is the substance that gives leaves their green color. 

✍ What Happens During Photosynthesis Step by Step?

Let’s understand what happens during photosynthesis step by step.

The plant needs to make Glucose or Food. For that they need two ingredients; Hydrogen and Carbon dioxide .

• First , Chlorophyll absorbs energy from sunlight.
• Second , roots transport water (H2O) to the leaves. [H2O is the molecular structure of the water. Two Hydrogen and One Oxygen.] Plants use sunlight energy to separate water molecules into hydrogen and oxygen.
• Third – Stomata receives Carbon dioxide (CO2) from the air.
• Fourth , After a series of chemical reactions, hydrogen and carbon dioxide combined together to produce glucose or food for the plant. After preparing food, there are veins, named Phloem , distributes glucose to all the parts of the plant.
• Fifth , oxygen is released into the air as a waste product, which becomes vital for us.

Chemical structure of photosynthesis

✍ Some Photosynthesis Facts 

FACT #1. The energy for photosynthesis comes from light.

FACT #2. During photosynthesis, plants use sunlight, water, carbon dioxide to make their food. 

FACT #3. Light energy is converted to chemical energy by chlorophyll.

FACT #4. The green color of leaves is due to chlorophyll.

FACT #5. Leaves change color in autumn because plants slow down the process of photosynthesis

FACT #6. Plants perform photosynthesis in organelles called chloroplasts. There are two parts of chloroplast i.e. grana and stroma

FACT #7. Only plants do not use photosynthesis. Rather some bacteria, such as cyanobacteria, and proteases, such as algae, are also producers. These single-celled organisms also contain chlorophyll.

FACT #8. All living things on this planet depend on organic molecules synthesized by plants as a result of photosynthesis.

In Conclusion,  

• Photosynthesis means “Photo” (means light) and “Synthesis” ( means putting together.)
• The process of making food by plant is called photosynthesis . 
• During photosynthesis, plants use sunlight, water, CO2 ( carbon dioxide i.e. present in air) to make their food. 
• The leaves are the food factory of the plant.
• Chlorophyll absorbs energy from sunlight.
• Stomata receives Carbon dioxide (CO2) from the air.
• After preparing food, Phloem distributes glucose (food) to all the parts of the plant.

I hope you understand the topic of photosynthesis for kids. Don’t forget to attempt a photosynthesis quiz for kids to check your knowledge. V isit here to know more about photosynthesis for kids.

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photosynthesis

[ foh-t uh - sin -th uh -sis ]

• the complex process by which carbon dioxide, water, and certain inorganic salts are converted into carbohydrates by green plants, algae, and certain bacteria, using energy from the sun and chlorophyll .

/ ˌfəʊtəʊsɪnˈθɛtɪk; ˌfəʊtəʊˈsɪnθɪsɪs /

• (in plants) the synthesis of organic compounds from carbon dioxide and water (with the release of oxygen) using light energy absorbed by chlorophyll
• the corresponding process in certain bacteria

/ fō′tō-sĭn ′ thĭ-sĭs /

• The process by which green plants, algae, diatoms, and certain forms of bacteria make carbohydrates from carbon dioxide and water in the presence of chlorophyll, using energy captured from sunlight by chlorophyll, and releasing excess oxygen as a byproduct. In plants and algae, photosynthesis takes place in organelles called chloroplasts . Photosynthesis is usually viewed as a two-step process. First, in the light reactions , the energy-providing molecule ATP is synthesized using light energy absorbed by chlorophyll and accessory pigments such as carotenoids and phycobilins, and water is broken apart into oxygen and a hydrogen ion, with the electron of the hydrogen transferred to another energy molecule, NADPH. The ATP and NADPH molecules power the second part of photosynthesis by the transfer of electrons. In these light-independent or dark reactions , carbon is broken away from carbon dioxide and combined with hydrogen via the Calvin cycle to create carbohydrates. Some of the carbohydrates, the sugars, can then be transported around the organism for immediate use; others, the starches, can be stored for later use.
• Compare chemosynthesis See Note at transpiration
• Use by green plants of the energy in sunlight to carry out chemical reactions , such as the conversion of carbon dioxide into oxygen . Photosynthesis also produces the sugars that feed the plant.

Discover More

Derived forms.

• ˌphotosynˈthetically , adverb
• photosynthetic , adjective

Other Words From

• pho·to·syn·thet·ic [ foh-t, uh, -sin-, thet, -ik ] , adjective
• photo·syn·theti·cal·ly adverb
• non·photo·syn·thetic adjective

Word History and Origins

Origin of photosynthesis 1

A Closer Look

Compare meanings.

How does photosynthesis compare to similar and commonly confused words? Explore the most common comparisons:

• chemosynthesis vs. photosynthesis
• photosynthesis vs. cellular respiration
• photorespiration vs. photosynthesis

Example Sentences

Specifically, he was interested in the protein-based "reaction centers" in spinach leaves that are the basic mechanism for photosynthesis—the chemical process by which plants convert carbon dioxide into oxygen and carbohydrates.

Algae and plants use photosynthesis to turn sunlight into food.

According to the Washington Post, this happens because as the days shorten and turn frigid, it’s not worth it for some trees to expend energy to conduct photosynthesis.

In a steady state, most of the energy captured by photosynthesis is used up by the furnace of respiration and metabolism burning on Earth’s surface by its infrared layer of life.

There’s no sunlight beneath half a mile of ice, so of course there’s no photosynthesis.

Nevertheless, it was required, and at least it was more fun than studying algebra or photosynthesis.

Re-solarizing the food chain should be our goal in every way—taking advantage of the everyday miracle that is photosynthesis.

As the microbes moved toward the light to carry out photosynthesis, they projected the image of the stencil.

Timiriazeff, in his Croonian Lecture, was the first to see the connexion between photosynthesis and the Lagado research.

On the other hand, their ancestors, the green or yellow mastigota, form new plasm by photosynthesis like true cells.

There the miracle of life consists merely of the chemical process of plasmodomism by photosynthesis.

Like von Baeyer's hypothesis, this assumes that formaldehyde and oxygen are the first products of photosynthesis.

In general, starch is the final product of photosynthesis in most green plants; but there are many exceptions to this.

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What is photosynthesis?

April 28, 2020 By Emma Vanstone Leave a Comment

Green plants make sugar for growth by a process called photosynthesis . Photosynthesis is a process where light energy is converted to chemical energy in the form of sugars. It’s a process that provides the main source of oxygen in the atmosphere and is essential for almost all life on Earth.

Photosynthesis made simple

Plants use sunlight, water, and carbon dioxide to create oxygen and energy in the form of glucose.

The glucose molecules created by photosynthesis act as fuel for cells and are used for cellular respiration and fermentation.

Photosynthesis Equation

Carbon dioxide + water  (and light ) ———> glucose and oxygen

Where does photosynthesis happen?

Photosynthesis takes place in chloroplast cells, which contain a substance called chlorophyll. It is chlorophyll, which gives plants their green colour.

Chloroplasts are one of the organelles in a plant cell. We made a jelly model of plant cell to learn about organelles and plant cell structure.

Sunlight is also needed to make chlorophyll. If plants are kept in the dark, they can’t make chlorophyll and will have yellow leaves! You can see this in our multicoloured cress caterpillar .

What affects the rate of photosynthesis

Four factors affect the rate of photosynthesis . The faster it occurs, the more the plant grows.

Light  – the more light there is, the faster photosynthesis occurs.

Water  – lack of water slows photosynthesis down.

Temperature  – photosynthesis works best at around 30 degrees Celsius.

Carbon dioxide levels  – photosynthesis is faster if there is more carbon dioxide in the air.

Plants make the energy to grow through a process called respiration . This uses the sugar produced by photosynthesis and oxygen.

How are leaves adapted for photosynthesis?

Leaves are green as they contain lots of chlorophyll, which absorbs sunlight.

They are thin and have a large surface area. This means they can absorb a lot of sunlight, and gases such as oxygen and carbon dioxide can pass in and out of the leaf easily.

Leaves have veins – xylem and phloem, which transport water, minerals and sugars around the plant.

Do only plants photosynthesise?

Plants, algae, and some types of bacteria use photosynthesis to create energy.

A large percentage of the Earth’s oxygen comes from phytoplankton in the oceans, which contain chlorophyll and use photosynthesis to create energy.

What is chlorophyll?

Chlorophyll is a green pigment which absorbs energy from blue and red light waves and reflects green light waves, which is why plants look green!

Chlorophyll in a plant is found in an organelle called a chloroplast. This is where photosynthesis occurs.

Why is photosynthesis important?

Photosynthesis also allows plants to make energy for growth and repair, and it has an important ecological impact.

Plants incorporate the carbon from carbon dioxide into organic molecules ( carbon fixation ). This creates a carbon source for animals who cannot create their own and also removes carbon dioxide from the air, slowing down the rate at which it builds up in the atmosphere.

Photosynthesis also creates oxygen, which is needed for most life on Earth!

Do you have more questions? You might find the answers in my collection of science questions for kids .

Last Updated on January 26, 2024 by Emma Vanstone

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